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- Viola brittoniana
Viola brittoniana — coast violet
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Facts
Viola brittoniana is a rare, acaulescent blue-flowered violet that occurs on the greater coastal plain in New England. It has distinctive leaves with narrow lobes and deep sinuses.
Habitat
Anthropogenic (man-made or disturbed habitats), bogs, floodplain (river or stream floodplains), forest edges, forests, marshes, meadows and fields, shores of rivers or lakes
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
Found this plant? Take a photo and post a sighting.
Characteristics
- Habitat
- terrestrial
- New England state
-
- Connecticut
- Massachusetts
- Flower petal color
- blue to purple
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf arrangement
- basal: the leaves are growing only at the base of the plant
- Leaf blade edges
- the edge of the leaf blade has lobes, or it has both teeth and lobes
- Flower symmetry
- there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Fusion of sepals and petals
- both the petals and sepals are separate and not fused
- Stamen number
-
- 1 or 2
- 5
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit length
- 10–18 mm
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Anther attachment
- the anther is attached by its base to the filament
- Anther opening
- the anthers have narrow slits or furrows that run lengthwise along the anthers
- Anther spurs
- there are spurs on the anthers
- Anther tube length
- 0 mm
- Calyx growth after flowering
- the calyx does not grow to cover or partially cover the fruit
- Calyx symmetry
- there is only one way to evenly divide the calyx (the calyx is bilaterally symmetrical)
- Carpels fused
- the carpels are fused to one another
- Cilia on petals
- the petal margins do not have cilia
- Cleistogamous flowers
-
- the plant has some cleistogamous flower
- there are no cleistogamous flowers on the plan
- Corolla morphology
- NA
- Corolla palate
- no
- Corona lobe length
- 0 mm
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Filament surface
- the filament is smooth, with no hairs or scales
- Flower appearance
-
- the flowers appear after the leaves have appeared
- the flowers appear at the same time as the leaves
- Flower description
- the flower has a superior ovary, and lacks a hypanthium
- Flower diameter
- 20–35 mm
- Flower length
- 13–22 mm
- Flower number
- 1
- Flower orientation
- the flower bends downwards or hangs downwards
- Flower petal color
- blue to purple
- Flower reproductive parts
- the flower has both pollen- and seed-producing parts
- Flower symmetry
- there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
- Flowers sunken into stem
- no
- Form of style
- the style is knob-like at the tip, and unbranched
- Fringed petal edges
- the petals are not fringed
- Fused stamen clusters
- NA
- Fusion of sepals and petals
- both the petals and sepals are separate and not fused
- Hairs on flower stalk
- the flower stalk has no hairs on it
- Horns in hoods (Asclepias)
- NA
- Hypanthium
- the flower does not have a hypanthium
- Hypanthium length
- 0 mm
- Inflorescence one-sided
- the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
- Inner tepals (Rumex)
- NA
- Interior flower disk
- the flower does not have an interior disc
- Length of peduncle
- Up to 150 mm
- Nectar spur
- the flower has one or more nectar spurs
- Number of branches in umbel
- 0
- Number of carpels
- 3
- Number of pistils
- 1
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Number of styles
- 1
- Ovary position
- the ovary is above the point of petal and/or sepal attachment
- Perianth shape
- NA
- Petal and sepal arrangement
- the flower includes two cycles of petal- or sepal-like structures
- Petal and sepal colors
- blue to purple
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal base
-
- the petal narrows abruptly at the base
- the petal narrows gradually or does not narrow at the base
- Petal folding in bud
- the petals in bud are arranged in a cycle with edges overlapping like roof shingles (imbricate)
- Petal folds or pleats
- the petals of the flower do not have folds or plaits
- Petal glandular dots or scales
- no
- Petal hairs (Viola)
- the side petals of the flower are hairy with hairs that are narrow or very slightly expanded at the tip, but not knob-shaped
- Petal hairs on inner/upper surface
- there are hairs on the inner/upper petal surface
- Petal length relative to sepals
- the petals are longer than the sepals
- Petal nectaries
- the petals do not have nectaries
- Petal number
- 5
- Petal shape
-
- the petal outline is oblong (rectangular, but with rounded ends)
- the petal outline is obovate (roughly egg-shaped, but with the widest point above the middle of the leaf blade)
- Petal tip shape
-
- the petal tip is retuse (with a blunt or rounded apex and a notch at the center)
- the petal tip is rounded
- Petal tips (Cuscuta)
- NA
- Raceme attachment (Veronica)
- NA
- Reproductive system
- all the flowers have both carpels and stamens (synoecious)
- Scales inside corolla
- no
- Sepal and petal color
- the sepals are different from the petals
- Sepal appearance
- the sepals are green or brown, and leaf-like in texture
- Sepal appendages
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Sepal auricles
- there is one auricle per sepal
- Sepal cilia
- the sepals do not have cilia
- Sepal color
- green to brown
- Sepal features
- NA
- Sepal length
- 6–10 mm
- Sepal number
- 5
- Sepal orientation
- the sepals are pressed against the corolla, or jutting stiffly upward
- Sepal relative length
- NA
- Sepal shape
- the sepal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)
- Sepal texture
- the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
- Sepal tip shape
-
- the sepal tip is acuminate (tapers to a very narrow point)
- the sepal tip is acute (is sharply pointed)
- Sepal uniformity
- all the sepals are about the same size
- Sepals fused only to sepals
- the sepals are separate from one another
- Spur length
- Up to 3.2 mm
- Spur number
- 1
- Stamen appendages
- stamen appendages are present
- Stamen attachment
-
- the stamens are not attached to the petals or tepals
- the stamens are not attached to the petals or tepals
- Stamen morphology
- the stamens within each cycle are the same
- Stamen number
-
- 1 or 2
- 5
- Stamen position relative to petals
- the stamens are lined up with the sepals
- Stamen relative length
- anything
- Stamens fused
- the stamens are not attached to one another
- Staminodes
- there are no staminodes on the flower
- Stigma position
- the stigmas are positioned at the tip of the style
- Style petal-like
- the styles are not petal-like
- Style relative length
- the stigma does not protrude beyond the mouth of the corolla
- Surface of ovary
- the ovary surface has no points, bumps or wrinkles
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene relative orientation
- NA
- Achene shape
- NA
- Achene surface (Polygonum)
- NA
- Achene type
- NA
- Berry color
- NA
- Capsule color (Viola)
- the capsule is entirely green
- Capsule ribs
- the capsule has no prominent ribs or wings
- Capsule splitting
- the capsule splits by three main valves, teeth or pores
- Fruit (pyxis) dehiscence
- NA
- Fruit beak length
- 0 mm
- Fruit cross-section
- the fruit is round in cross-section
- Fruit features (Brassicaceae)
- NA
- Fruit length
- 10–18 mm
- Fruit length relative to sepals
- the fruit is longer than its associated sepals
- Fruit locules
- three
- Fruit shape
- the fruit is ovoid (egg-shaped)
- Fruit stalk orientation
- the fruits curve or droop downwards
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit type (specific)
- the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)
- Hair type on fruit
- the hairs on the fruits are simple (not branched), don’t have glands, and are not woolly
- Hairs on fruit
- the fruits are not hairy
- Legumes (Fabaceae)
- NA
- Mericarp length
- 0 mm
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
- NA
- Ovary stipe
- the ovary or fruit does not have a stipe
- Ovary stipe length
- 0 mm
- Placenta arrangement
- the plant has parietal placentation, where ovules develop on the wall or slight outgrowths of the wall forming broken partitions within a compound ovary
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- NA
- Schizocarpic fruit segments
- 0
- Seed length
- 1.8–2 mm
- Seed relative length
- the seed is longer than it is wide
- Seed surface
- the seed is smooth or without clear markings
- Seeds comose
- no hairs
- Septum in fruit (Brassicaceae)
- NA
- Wings on fruit
- the fruit does not have wings on it
- prickles on fruits
- the fruits do not have thorn-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades do not have glandular dots or scales
- Sap
- the sap is clear and watery
- Sap color
- the sap is clear
-
Growth form
- Growth form
- the plant is an herb (it has self-supporting stems)
- Horizontal rooting stem
- the plant does not have stolons
- Lifespan
- the plant lives more than two years
- Parasitism
- the plant is not parasitic
- Plant color
- the leaves or young stems of the plant are green
- Plants darken when dry
- no
- Spines on plant
- the plant has no spines
- Underground organs
- the plant has a rhizome (a horizontal underground stem with roots growing from it)
-
Leaves
- Bracteole number (Apiaceae)
- 0
- Bracteoles
- the plant has bracteoles between the primary bracts and the flowers
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length (compound lvs only)
- 0 mm
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Final leaf segment width (compound lvs only)
- 0 mm
- Floral bracts
- the flower has one or more bracts associated with it
- Hairs on leaf stalk
- the petiole has no hairs on it
- Hairs on underside of leaf
- the underside of the leaf is not hairy, or it has very few hairs
- Hairs on upper side of leaf
- the upper side of the leaf is not hairy, or it has very few hairs
- Hooked hairs on underside of leaf
- no
- Inflated hairs on leaf
- the leaf blade does not have inflated hairs on it
- Leaf arrangement
- basal: the leaves are growing only at the base of the plant
- Leaf blade base
- the leaf has a distinct leaf stalk (petiole)
- Leaf blade base shape
- the base of the leaf blade is cordate (heart-shaped, has rounded lobes at the base)
- Leaf blade base symmetry
- the leaf blade base is symmetrical
- Leaf blade bloom
- the underside of the leaf has no noticeable bloom
- Leaf blade edges
- the edge of the leaf blade has lobes, or it has both teeth and lobes
- Leaf blade flatness
- the leaf is flat (planar) at the edges
- Leaf blade hairs
- NA
- Leaf blade length
- 40–90 mm
- Leaf blade shape
-
- the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
- the leaf blade is reniform (kidney-shaped; wider than long)
- the leaf blade is triangular, with the stalk or attachment point on one of the sides
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are green, with an expanded blade and a leaf-like texture
- Leaf hair orientation
- NA
- Leaf sheath length
- 0 mm
- Leaf shiny
- the upper side of the leaf is dull or slightly shiny
- Leaf spines
- there are no spines on the leaf edges
- Leaf stalk
- the leaves have leaf stalks
- Leaf stalk attachment to leaf
- the petiole attaches at the basal margin of the leaf blade
- Leaf stalk base
- the petiole base is narrow where it attaches to the stem
- Leaf stalk length
- Up to 220 mm
- Leaf teeth and lobes
-
- the leaf blade margin has outward-pointing teeth
- the leaf has lobes that radiate from the base, somewhat like a hand
- Leaf tip
- the tip of the leaf blade is acute (sharply pointed)
- Leaf tufts in axils
-
- NA
- there are no clusters of smaller leaves growing out of axils
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf types
- NA
- Leaf variation
- NA
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
- the leaves are growing only at the base of the plant (i.e., basal)
- Pinnately compound leaf type
- NA
- Specific leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Stipels
- NA
- Stipule edges
-
- the stipule margins do not have teeth
- the stipule margins have teeth on them
- Stipule features
- NA
- Stipule fused to leaf stalk
- the stipules are not fused to the petioles
- Stipule length
- Up to 20 mm
- Stipule shape
- the stipules are linear (very narrow with more or less parallel sides)
- Stipules
- the plant has stipules
-
Place
- Habitat
- terrestrial
- New England state
-
- Connecticut
- Massachusetts
- Specific habitat
-
- bogs
- edges of forests
- forests
- man-made or disturbed habitats
- marshes
- meadows or fields
- river or stream floodplains
- shores of rivers or lakes
-
Scent
- Plant odor
- the plant does not have much of a smell
-
Stem, shoot, branch
- Branched tendrils
- NA
- Direction of stem hairs
- NA
- Flowering stem cross-section
- NA
- Hair between stem nodes
- NA
- Hairs between stem nodes
- NA
- Hooked hairs on stem between nodes
- NA
- Leaves on stem
- the flowering stem has no leaves above the base
- Length of hairs between stem nodes
- 0 mm
- Plant height
- 2–45 cm
- Stem bloom
- NA
- Stem hair distribution
- NA
- Stem nodes swollen
- NA
- Stem orientation
- NA
- Stem roughness between nodes
- NA
- Stem spacing
- NA
- Stem succulence
- NA
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
- Wings on stem
- NA
Wetland status
Occurs in wetlands or non-wetlands. (Wetland indicator code: FAC)
In New England
Distribution
- Connecticut
- present
- Maine
- absent
- Massachusetts
- present
- New Hampshire
- absent
- Rhode Island
- absent
- Vermont
- absent
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Connecticut
- extremely rare (S-rank: S1), endangered (code: E)
- Maine
- historical (S-rank: SH), potentially extirpated (code: PE)
- Massachusetts
- rare (S-rank: S2), threatened (code: T)
From Flora Novae Angliae dichotomous key
5. Viola brittoniana Pollard NC
coast violet. Viola atlantica Britt.; V. pedatifida G. Don ssp. brittoniana (Pollard) McKinney • CT, MA. Fields, meadows, trail edges, and forest clearings adjacent to rivers and coastal marshes, also peaty river shores. Previous reports of this species in ME were based on collections of a hybrid between Viola brittoniana and V. sororia (specimens at GH!, NEBC!).
5×7. Viola brittoniana × Viola cucullata → Viola ×notabilis Bickn. is a very rare violet hybrid known from CT, MA, ME (the ME population is an escape from cultivation, the others are naturally occurring hybrids in wild populations). This hybrid is recognized by its ± glabrous herbage, robust size (characteristic of many V. cucullata hybrids), its variably lobed leaves with lobes broader and shorter than in V. brittoniana, and its longer, more cylindrical hairs on the lateral petals (not conspicuously knob-shaped as in V. cucullata).
5×11. Viola brittoniana × Viola lanceolata → This very rare violet hybrid is known from MA. It is relatively easy to distinguish due to the very different morphologies of the parental species. It is a relatively glabrous plant with cyanic corollas, short stolons later in the season, broad-lanceolate to narrow-ovate leaf blades that are lobed with an elongate central lobe and short lateral lobes (rather than triangular-ovate to suborbicular and with elongate lobes nearly as long as the entire blade in V. brittoniana).
5×18. Viola brittoniana × Viola pectinata → This very rare violet hybrid is known from CT, MA. It has lobed leaf blades somewhat similar to Viola brittoniana except that the lobes are broader relative to length and the sinuses are not as deep. Further, the early leaves are usually less lobed relative to the later leaves. Given the floral and fruiting similarity of V. brittoniana and V. pectinata, the flowers appear as in both taxa (i.e., there is no altering of the reproductive morphology as there would be in other hybrids; e.g., shorter and broader tipped petal hairs in hybrids with V. cucullata, relatively broader sepals with some cilia in hybrids with V. sororia).
5×25a. Viola ×mulfordae Pollard is a very rare violet hybrid known from CT, MA. It resembles V. brittoniana in its conspicuous lobed leaf blades, but the lobes are shorter and broader than in V. brittoniana and often with an elongate central lobe (contributing to the longer-than-wide aspect of the leaf blade). Further, the petioles and leaf blades are pubescent (glabrous in V. brittoniana).
5×27. Viola brittoniana × Viola sororia → Viola ×insolita House is a very rare violet hybrid known from CT, ME. It has variably lobed leaves (with broader and shorter segments than in V. brittoniana) with a prominent central lobe that are at least sparsely pubescent, sepals with cilia (at least on the auricles), prostrate to ascending cleistogamous peduncles, green or purple-spotted capsules, and brown seeds (pale brown to orange-yellow in V. brittoniana, light yellow-brown to brown or dark brown in V. sororia).
Native to North America?
Yes
Sometimes confused with
- Viola palmata:
- sepals ovate, rounded to obtuse at the apex, usually at least sparsely ciliate, capsules green and spotted or suffused with purple, and plant heterophyllous, the early leaves unlobed and the later leaves lobed (vs. V. brittoniana, with sepals lanceolate, acuminate at the apex, eciliate, capsules green, and plants homophyllous, all the leaves lobed).
- Viola subsinuata:
- sepals ovate, rounded to obtuse at the apex, usually at least sparsely ciliate, capsules green and spotted or suffused with purple, and leaf blades usually pubescent(vs. V. brittoniana, with sepals lanceolate, acuminate at the apex, eciliate, capsules green, and leaf blades essentially glabrous).
Synonyms
- Viola atlantica Britt.
- Viola pedatifida G. Don ssp. brittoniana (Pollard) McKinney