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- Ilex glabra
Ilex glabra — evergreen winterberry, inkberry
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Facts
Inkberry is a widely planted shrub with glossy dark green leaves and a rounded growth form. Its dark black, "inky" berries persist well into winter; birds disperse the seeds. Although it ranges broadly throughout the Atlantic coastal plain, native populations are uncommon in New England. This species grows well in a variety of conditions, from wetlands (where it is sometimes used to mitigate or replicate these habitats) to dunes. It is a popular choice for screen-plantings in the garden.
Habitat
Bogs, shores of rivers or lakes, swamps
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
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Characteristics
- Habitat
-
- terrestrial
- wetlands
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Growth form
- the plant is a shrub (i.e., a woody plant with several stems growing from the base)
- Leaf type
- the leaf blade is simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaves per node
- there is one leaf per node along the stem
- Leaf blade edges
- the edge of the leaf blade has teeth
- Leaf duration
- the leaves remain green all winter
- armature on plant
- the plant does not have spines, prickles, or thorns
- Leaf blade length
- 15–50 mm
- Leaf blade width
- 7–20 mm
- Leaf stalk
- the leaves have leaf stalks
- Fruit type (general)
- the fruit is fleshy
- Bark texture
- the bark of an adult plant is thin and smooth
- Twig winter color
-
- brown
- gray
- Bud scale number
- there are three or more scales on the winter bud, and they overlap like shingles, with one edge covered and the other edge exposed
-
Buds or leaf scars
- Bud position relative to scar
- the winter bud has a leaf scar just below it
- Bud scale number
- there are three or more scales on the winter bud, and they overlap like shingles, with one edge covered and the other edge exposed
- Bud scar shape (Fraxinus)
- NA
- Collateral buds
- there are no collateral buds on the sides of the branches
- Leaf scar arrangement
- there is one leaf scar per node on the stem or twig
- Superposed buds
- there are no superposed buds on the branch
- Winter bud scales
- the winter bud is perulate (partially or completely covered with one or more scales)
- Winter bud shape
- the winter buds are ovoid (egg-shaped)
- Winter bud stalks
- the winter buds have no stalks
-
Flowers
- Anther color
- the anthers show no hint of a pink, reddish or purplish tint
- Carpels fused
- the carpels are fused to one another
- Enlarged sterile flowers
- there are no enlarged sterile flowers on the plant
- Flower petal color
- white
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Hairs on ovary (Amelanchier)
- NA
- Hypanthium present
- the flower does not have a hypanthium
- Inflorescence hairs
- there are no hairs on the inflorescence
- Inflorescence position
- the inflorescences grow on the twigs
- Inflorescence type
-
- the flowers grow out of the axil (point where a branch or leaf is attached to the main stem)
- the inflorescence differs from the choices given
- the inflorescence has only one flower on it
- Number of pistils
- 1
- Ovary position
- the ovary is above the point of petal and/or sepal attachment
- Petal and sepal arrangement
- the flower includes two cycles of petal- or sepal-like structures
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal fusion
- the perianth parts are fused to form a tube, cup, or bell shape
- Sepal appearance
- the sepals resemble leaves in color and texture
- Sepal tip glands
- there are no glands at the tips of the sepal lobes
- Sepals fused only to sepals
- the sepals are fused to each other (not other flower parts), at least near their bases
- Stamen number
-
- 4
- 5
- Stamen position relative to petals
- the stamens are lined up with the petals (antepetalous)
- Stamens fused
- the stamens are not fused to one another
-
Fruits or seeds
- Berry color
- black
- Fruit tissue origin
- there are no flower parts that form part of the fruit
- Fruit type (general)
- the fruit is fleshy
- Fruit type (specific)
- the fruit is a drupe (fleshy, with a firm inner ovary wall that encloses a single seed)
- Nut with spines (Fagaceae)
- NA
- Wings on fruit
- there are no wings on the fruit
-
Glands or sap
- Sap color
- the sap is clear and watery
- Stalked glands on fruit (Rosa)
- NA
-
Growth form
- Growth form
- the plant is a shrub (i.e., a woody plant with several stems growing from the base)
-
Leaves
- Hairs on underside of leaf blade
- the underside of the leaf has no hairs
- Hairs on upper side of leaf blade
- the upper side of the leaf is not hairy, or has very few hairs
- Leaf blade base shape
- the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
- Leaf blade base symmetry
- the leaf blade base is symmetrical
- Leaf blade bloom
- the underside of the leaf has no noticeable bloom
- Leaf blade edges
- the edge of the leaf blade has teeth
- Leaf blade edges (Acer)
- NA
- Leaf blade flatness
- the leaf is flat (planar) at the edges
- Leaf blade hairs
- NA
- Leaf blade length
- 15–50 mm
- Leaf blade scales
- there are scales on the leaf blades
- Leaf blade shape
-
- the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
- the leaf blade is oblanceolate (lance-shaped, but with the widest point above the middle of the leaf blade)
- the leaf blade is oblong (rectangular but with rounded ends)
- Leaf blade texture
- the leaf blade is coriaceous (has a firm, leathery texture)
- Leaf blade translucent dots
- there are no translucent dots on the leaf blade
- Leaf blade vein pattern
- the main veins of the leaf blade are pinnate (the secondary veins branch off at intervals from the main central vein) and non-arcuate (not arched towards the leaf tip)
- Leaf blade veins
- the leaf blade has one main vein running from the base toward the tip
- Leaf blade width
- 7–20 mm
- Leaf duration
- the leaves remain green all winter
- Leaf form
- the plant is broad-leaved (with broadly flattened leaf blades)
- Leaf lobe tips (Quercus)
- NA
- Leaf midrib glands
- the midrib of the leaf blade lacks glands on the upper surface
- Leaf stalk
- the leaves have leaf stalks
- Leaf stalk attachment to leaf
- the petiole attaches at the basal margin of the leaf blade
- Leaf stalk nectaries
- there are no nectaries on the leaf stalk
- Leaf stalk shape
- the leaf stalk is not flattened
- Leaf teeth
-
- the leaf blade margin is crenate (with rounded teeth) or crenulate (with tiny, rounded teeth)
- the leaf blade margin is serrate (with forward-pointing) or dentate (with outward-pointing) with medium-sized to coarse teeth
- Leaf teeth hairs (Carya)
- NA
- Leaf type
- the leaf blade is simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaves per node
- there is one leaf per node along the stem
- Specific leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets
- Stipules
- the plant has stipules
-
Place
- Habitat
-
- terrestrial
- wetlands
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Specific habitat
-
- bogs
- shores of rivers or lakes
- swamps
-
Scent
- Plant odor
- the plant does not have much of an odor, or it has an unpleasant or repellant odor
-
Stem, shoot, branch
- Aerial roots
- the plant has no aerial roots
- Bark texture
- the bark of an adult plant is thin and smooth
- Branch brittleness (willows only)
- NA
- Branch cross-section
- the branch is circular in cross-section, or it has five or more sides, so that there are no sharp angles
- First-year cane (Rubus)
- NA
- Lenticels on twigs
- there are no lenticels on the twigs, or they are very hard to see
- Pith shape
- the outline of the pith in a twig is roughly round
- Pith type
- the pith inside the twig is solid, completely filled with spongy tissue
- Twig bloom
- there is no bloom on the twig
- Twig hairs
- the twigs have hairs, but the hairs do not have glands
- Twig papillae (Vaccinium species only)
- NA
- Twig scales
- there are no scales on the twig surface
- Twig winter color
-
- brown
- gray
- Wings on branch
- the branch does not have wings on it
- armature on plant
- the plant does not have spines, prickles, or thorns
Wetland status
Usually occurs in wetlands, but occasionally in non-wetlands. (Wetland indicator code: FACW)
In New England
Distribution
- Connecticut
- present
- Maine
- present
- Massachusetts
- present
- New Hampshire
- present
- Rhode Island
- present
- Vermont
- absent
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Connecticut
- rare (S-rank: S2), threatened (code: T)
- Maine
- extremely rare (S-rank: S1), endangered (code: E)
- Massachusetts
- fairly widespread (S-rank: S4)
- New Hampshire
- historical (S-rank: SH), endangered (code: E)
From Flora Novae Angliae dichotomous key
3. Ilex glabra (L.) Gray N
evergreen winterberry. CT, MA, ME, NH, RI. Swamps, lake shores, bogs.
Native to North America?
Yes
Sometimes confused with
- Ilex crenata:
- leaf blades with mostly 4–10 teeth per margin, the teeth usually not restricted to the apical portion of the blade, and flowers 4-merous (vs. I. glabra, with leaf blades with 1–3 teeth per margin, the teeth usually restricted to the apical 1/3 of the blade, and flowers 6- to 8-merous).