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- Hackelia deflexa
Hackelia deflexa — nodding stickseed
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Facts
Nodding stickseed has a wide distribution in northern North America, but in New England it is very rare. It inhabits open, dry, calcareous habitats including rocky forests and cliff bases. Despite two new populations being found in Maine in 2000, this plant appears to be in decline in New England, chiefly due to loss of habitat in Vermont's Champlain Valley.
Habitat
Cliffs, balds, or ledges, forests, talus and rocky slopes
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
Found this plant? Take a photo and post a sighting.
Characteristics
- Habitat
- terrestrial
- New England state
-
- Maine
- New Hampshire
- Vermont
- Flower petal color
-
- blue to purple
- white
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf arrangement
- alternate: there is one leaf per node along the stem
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Stamen number
- 5
- Fruit type (general)
- the fruit is dry but does not split open when ripe
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Anther opening
- the anthers have narrow slits or furrows that run lengthwise along the anthers
- Calyx symmetry
- there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
- Carpels fused
- the carpels are fused to one another
- Cleistogamous flowers
- there are no cleistogamous flowers on the plan
- Corolla palate
- no
- Corona lobe length
- 0 mm
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Flower description
- the flower has a superior ovary, and lacks a hypanthium
- Flower petal color
-
- blue to purple
- white
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Flowers sunken into stem
- no
- Fused stamen clusters
- NA
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Horns in hoods (Asclepias)
- NA
- Hypanthium
- the flower does not have a hypanthium
- Inflorescence one-sided
- the flowers are all or nearly all arrayed on one side of the inflorescence axis or branches of the inflorescence
- Inner tepals (Rumex)
- NA
- Nectar spur
- the flower has no nectar spurs
- Number of pistils
- 1
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Number of styles
- 1
- Ovary position
- the ovary is above the point of petal and/or sepal attachment
- Petal and sepal arrangement
- the flower includes two cycles of petal- or sepal-like structures
- Petal and sepal colors
-
- blue to purple
- white
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal folds or pleats
- the petals of the flower do not have folds or plaits
- Petal hairs (Viola)
- NA
- Petal number
- 5
- Petal tips (Cuscuta)
- NA
- Reproductive system
- all the flowers have both carpels and stamens (synoecious)
- Sepal and petal color
- the sepals are different from the petals
- Sepal appendages
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Sepal number
- 5
- Stamen attachment
-
- the stamens are attached at or near the bases of the petals or tepals
- the stamens are attached at or near the bases of the petals or tepals
- Stamen number
- 5
- Stamen position relative to petals
- the stamens are lined up with the sepals
- Staminodes
- there are no staminodes on the flower
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene relative orientation
- NA
- Achene shape
- NA
- Achene surface (Polygonum)
- NA
- Achene type
- NA
- Berry color
- NA
- Capsule color (Viola)
- NA
- Capsule ribs
- NA
- Capsule splitting
- NA
- Fruit (pyxis) dehiscence
- NA
- Fruit beak length
- 0 mm
- Fruit features (Brassicaceae)
- NA
- Fruit locules
- four
- Fruit type (general)
- the fruit is dry but does not split open when ripe
- Fruit type (specific)
- the fruit is a schizocarp (when dry it splits into sections, each holding one or more seeds)
- Legumes (Fabaceae)
- NA
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
- NA
- Ovary stipe
- the ovary or fruit does not have a stipe
- Placenta arrangement
- the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- the fruit is not flattened
- Schizocarpic fruit segments
- 4
- Septum in fruit (Brassicaceae)
- NA
- Wings on fruit
- the fruit does not have wings on it
- prickles on fruits
- the fruits are armed with thorns-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades do not have glandular dots or scales
- Sap
- the sap is clear and watery
- Sap color
- the sap is clear
-
Growth form
- Horizontal rooting stem
- the plant does not have stolons
- Lifespan
- the plant lives for two years
- Parasitism
- the plant is not parasitic
- Plant color
- the leaves or young stems of the plant are green
- Plants darken when dry
- no
- Spines on plant
- the plant has no spines
-
Leaves
- Bracteole number (Apiaceae)
- 0
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length (compound lvs only)
- 0 mm
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Final leaf segment width (compound lvs only)
- 0 mm
- Hairs on underside of leaf
- the underside of the leaf is fuzzy or hairy
- Leaf arrangement
- alternate: there is one leaf per node along the stem
- Leaf blade base
-
- the leaf has a distinct leaf stalk (petiole)
- the leaf has no stalk
- Leaf blade base shape
- the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf blade length
- At least 10 mm
- Leaf blade shape
-
- the leaf blade is elliptic (widest near the middle and tapering at both ends)
- the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
- the leaf blade is oblanceolate (lance-shaped, but with the widest point above the middle of the leaf blade)
- the leaf blade is oblong (rectangular but with rounded ends)
- the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are green, with an expanded blade and a leaf-like texture
- Leaf spines
- there are no spines on the leaf edges
- Leaf teeth and lobes
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf types
- There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
- there is one leaf per node along the stem
- Pinnately compound leaf type
- NA
- Specific leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Stipels
- NA
- Stipule features
- NA
- Stipules
- there are no stipules on the plant
-
Place
- Habitat
- terrestrial
- New England state
-
- Maine
- New Hampshire
- Vermont
- Specific habitat
-
- cliffs, balds, or ledges
- forests
- talus or rocky slopes
-
Scent
- Plant odor
- the plant does not have much of a smell
-
Stem, shoot, branch
- Branched tendrils
- NA
- Flowering stem cross-section
- the flowering stem is circular, or with lots of small angles so that it is roughly circular
- Hairs between stem nodes
- the hairs on the stem are plain, without glands or branches, and not tangled
- Leaves on stem
- there is at least one full leaf above the base of the flowering stem
- Plant height
- Up to 150 cm
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
Wetland status
Not classified
In New England
Distribution
- Connecticut
- absent
- Maine
- present
- Massachusetts
- absent
- New Hampshire
- present
- Rhode Island
- absent
- Vermont
- present
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Maine
- extremely rare (S-rank: S1)
ssp. americana
- Maine
- extremely rare (S-rank: S1), endangered (code: E)
- New Hampshire
- historical (S-rank: SH), endangered (code: E)
- Vermont
- rare (S-rank: S2), threatened (code: T)
Subspecies and varieties
Our subspecies is Hackelia deflexa (Wahlenb.) Opiz ssp. americana.
From Flora Novae Angliae dichotomous key
1. Hackelia deflexa (Wahlenb.) Opiz ssp. americana (Gray) A. & D. Löve NC
nodding stickseed. Hackelia americana (Gray) Fern; H. deflexa (Wahlenb.) Opiz var. americana (Gray) Fern. & I.M. Johnston; Lappula americana (Gray) Rydb.; L. deflexa (Gray) Rydb. ssp. americana (Gray) Hultén; L. deflexa (Gray) Rydb. var. americana (Gray) Greene • ME, NH, VT. Rocky forests and cliff bases, in regions of high-pH bedrock.
Native to North America?
Yes
Sometimes confused with
- Hackelia virginiana:
- mericarps with 10–25 glochids on the outer surface that are of similar length to the marginal glochids and basal leaf blades broad-ovate to cordate-ovate (vs. H. deflexa, with mericarps with a single row of glochids on each margin, sometimes with 1–3 small glochids on the outer surface, and basal leaf blades narrow-ovate to oblanceolate).
Synonyms
- Hackelia americana (Gray) Fern
- Hackelia deflexa (Wahlenb.) Opiz var. americana (Gray) Fern. & I.M. Johnston
- Lappula americana (Gray) Rydb.
- Lappula deflexa (Gray) Rydb. ssp. americana (Gray) Hultén
- Lappula deflexa (Gray) Rydb. var. americana (Gray) Greene