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Botrychium angustisegmentum — narrow triangle moonwort

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Facts

Narrow triangle moonwort has a dark green, fleshy blade that appears in late spring or early summer and can persist into October. This species releases its spores later than most moonworts.

Habitat

Bogs, fens, forests, meadows and fields, swamps, wetland margins (edges of wetlands)

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
  • terrestrial
  • wetlands
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Leaf divisions
  • the leaf blade is compound (divided into leaflets)
  • the leaf blade is lobed
  • the leaf blade is twice compound (divided into leaflets, which are further divided into leaflets)
Plant growth form
the leaves grow on a stalk that also holds the sporophore, or spore-bearing leaf
Spore-bearing leaflets
the spore-bearing fronds are dramatically different from the sterile fronds
Sorus shape
there are no sori, or they are concealed in leaf segments or hardened, capsule-like structures derived from a modified leaflet
Leaf stalk scales
there are no scales on the leaf stalk
Leaf stalk hairs
there are no hairs on the leaf stalk
Leaf blade length
1–5 cm
Leaf vein tips
the veins go all the way to the edge of the leaf blade
Show all characteristics
  • Growth form

    Life form
    the plant is herbaceous and terrestrial
    Life stage
    the plant is visible as a typical leaf-bearing fern (sporophyte)
    Spore-bearing leaflets
    the spore-bearing fronds are dramatically different from the sterile fronds
  • Leaves

    Features of leaves
    there are no special features on the leaves
    Leaf blade length
    1–5 cm
    Leaf blade shape
    the leaf blades are roughly triangular
    Leaf blade tip shape
    the tip of the leaf blade is a sharp point (acute)
    Leaf blade width
    At least 1.5 cm
    Leaf divisions
    • the leaf blade is compound (divided into leaflets)
    • the leaf blade is lobed
    • the leaf blade is twice compound (divided into leaflets, which are further divided into leaflets)
    Leaf lifespan
    the leaves drop off in winter
    Leaf stalk color
    • green
    • yellow to brown
    Leaf stalk hairs
    there are no hairs on the leaf stalk
    Leaf stalk length
    60–200 mm
    Leaf stalk relative length
    the leaf stalk is more than three quarters as long as the blade
    Leaf stalk scale location
    there are no scales on the leaf stalk
    Leaf stalk scales
    there are no scales on the leaf stalk
    Leaf stalk vessels
    2 bundles
    Leaf vein branching
    the secondary veins of the leaf blade branch dichotomously (two equal branches at each branch point)
    Leaf vein tips
    the veins go all the way to the edge of the leaf blade
    Leaflet relative size
    • the bottom leaflets are about half as long as, to slightly longer than, the leaflets from the middle of the frond
    • the bottom leaflets are at least one and a half times as long as the leaflets from the middle of the frond
    Leaflet stalks
    • the leaflets are stalked
    • the leaflets do not have stalks
    Lobe or leaflet length
    4–33 mm
    Lobe or leaflet pairs
    1–8
    Lobe or leaflet shape
    • the lobe or leaflet is extremely narrow, thread-like
    • the lobe or leaflet is rectangular but with rounded ends (oblong)
    Lobe or leaflet width
    1–2 mm
    Plant growth form
    the leaves grow on a stalk that also holds the sporophore, or spore-bearing leaf
    final leaf segment margin
    the topmost lobe or leaflet of the leaf blade has a smooth or lobed edge
  • Place

    Habitat
    • terrestrial
    • wetlands
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • bogs
    • edges of wetlands
    • fens
    • forests
    • meadows or fields
    • swamps
  • Spores or spore cones

    Sorus features
    there are no special features on the sorus
    Sorus shape
    there are no sori, or they are concealed in leaf segments or hardened, capsule-like structures derived from a modified leaflet
    Sporangia location
    the spores are formed on a stalk (sporangiophore)
    Sporangium type
    the spores are enclosed in a hardened, capsule-like structure (sporocarp)
    Spore forms
    there is only one type of spore present

Wetland status

Usually occurs in wetlands, but occasionally in non-wetlands. (Wetland indicator code: FACW)

In New England

Distribution

Connecticut
absent
Maine
absent
Massachusetts
absent
New Hampshire
absent
Rhode Island
absent
Vermont
absent

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

Massachusetts
rare to uncommon (S-rank: S2S3)
Rhode Island
extremely rare (S-rank: S1), concern (code: C)
Vermont
fairly widespread (S-rank: S4)

From Flora Novae Angliae dichotomous key

1.  Botrychium angustisegmentum (Pease & Moore) Fern. N

narrow triangle moonwort. Botrychium lanceolatum (Gmel.) Angst. ssp. angustisegmentum (Pease & Moore) Clausen; B. lanceolatum (Gmel.) Angst. var. angustisegmentum Pease & Moore • CT, MA, ME, NH, RI, VT. Forests, meadows, swamp margins, and peaty openings.

Native to North America?

Yes

Sometimes confused with

Botrychium matricariifolium:
vegetative portion of leaf oblong to ovate and usually short-stalked (vs. B. angustisegmentum, with vegetative portion of leaf +/- triangular and unstalked).

Synonyms

  • Botrychium lanceolatum (Gmel.) Angst. ssp. angustisegmentum (Pease & Moore) Clausen
  • Botrychium lanceolatum (Gmel.) Angst. var. angustisegmentum Pease & Moore

Genus

Botrychium