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Native Plant Trust: Go Botany Discover thousands of New England plants

Verbena simplex — narrow-leaved vervain

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Facts

Narrow-leaved vervain is widespread throughout the eastern half of North America, but very rare and in decline in New England. Perhaps a tenth of the recorded locations still harbor the plant, which grows on open, dry habitats such as fields, sandplains and road cuts. Populations have disappeared from sites subject to competition with other plants, or canopy closure due to natural succession.

Habitat

Anthropogenic (man-made or disturbed habitats), meadows and fields, ridges or ledges, sandplains and barrens

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
terrestrial
New England state
  • Connecticut
  • Massachusetts
  • Vermont
Flower petal color
blue to purple
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
opposite: there are two leaves per node along the stem
Leaf blade edges
the edge of the leaf blade has teeth
Flower symmetry
  • there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
  • there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
Number of sepals, petals or tepals
there are five petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
4
Fruit type (general)
the fruit is dry and splits open when ripe
Fruit length
2.4–2.8 mm
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Anther attachment
    the anther is attached near its midpoint to the filament
    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Anther spurs
    the anthers do not have spurs on them
    Anther tube length
    0 mm
    Calyx growth after flowering
    the calyx does not grow to cover or partially cover the fruit
    Calyx symmetry
    • there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    • there is only one way to evenly divide the calyx (the calyx is bilaterally symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cilia on petals
    the petal margins do not have cilia
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla morphology
    the flower has two prominent lips
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower orientation
    the flower points upwards or is angled outwards
    Flower petal color
    blue to purple
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    • there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
    • there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is lobed at the tip, and unbranched
    Fringed petal edges
    the petals are not fringed
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Hairs on flower stalk
    NA
    Hairs on inflorescence
    the axis of the inflorescence has hairs entirely without glands
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Hypanthium length
    0 mm
    Inflorescence length
    50–200 mm
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inflorescence width
    5–10 mm
    Inner tepals (Rumex)
    NA
    Length of flower stalk
    0 mm
    Marks on petals
    there are no noticeable marks on the petals
    Nectar spur
    the flower has no nectar spurs
    Number of branches in umbel
    0
    Number of carpels
    2
    Number of pistils
    1
    Number of sepals, petals or tepals
    there are five petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Perianth shape
    • the perianth is funnelform (funnel-shaped)
    • the perianth is salverform (with a slender basal tube and a more-or-less horizontally spreading mouth)
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    blue to purple
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal base
    the petal narrows gradually or does not narrow at the base
    Petal folding in bud
    • the petals are arranged some other way in bud
    • the petals in bud are arranged in a cycle with edges overlapping like roof shingles (imbricate)
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal glandular dots or scales
    no
    Petal hairs (Viola)
    NA
    Petal hairs on inner/upper surface
    there are hairs on the inner/upper petal surface
    Petal length
    2–4 mm
    Petal length relative to sepals
    the petals are longer than the sepals
    Petal number
    5
    Petal shape
    • the petal outline is oblong (rectangular, but with rounded ends)
    • the petal outline is obovate (roughly egg-shaped, but with the widest point above the middle of the leaf blade)
    Petal tip shape
    • the petal tip is lobed or fringed
    • the petal tip is obtuse (bluntly pointed)
    • the petal tip is retuse (with a blunt or rounded apex and a notch at the center)
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appearance
    the sepals are green or brown, and leaf-like in texture
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal auricles
    the sepals have no auricles
    Sepal color
    green to brown
    Sepal length
    2–5 mm
    Sepal number
    5
    Sepal orientation
    the sepals are pressed against the corolla, or jutting stiffly upward
    Sepal relative length
    the sepal lobes are shorter than the fused portion
    Sepal shape
    • the sepal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)
    • the sepal outline is linear (extremely narrow, thread-like)
    Sepal texture
    the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
    Sepal tip shape
    the sepal tip is lobed or fringed
    Sepal uniformity
    one or more of the sepals is much narrower or shorter than the others
    Sepals fused only to sepals
    the sepals are fused to each other (not other flower parts), at least near their bases
    Spur length
    0 mm
    Spur number
    NA
    Stamen appendages
    stamen appendages are absent
    Stamen attachment
    • the stamens are attached at or near the bases of the petals or tepals
    • the stamens are attached at or near the bases of the petals or tepals
    Stamen lengths differ
    the stamens are didynamous (two long stamens and two short ones)
    Stamen morphology
    the stamens within a cycle differ in length or width
    Stamen number
    4
    Stamen position relative to petals
    the stamens are lined up with the sepals
    Stamen relative length
    anything
    Stamens fused
    the stamens are not attached to one another
    Stigma position
    the stigmas are positioned at the tip of the style
    Style petal-like
    the styles are not petal-like
    Style relative length
    the stigma does not protrude beyond the mouth of the corolla
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    the upper lip of the bilabiate corolla has four or more lobes

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    NA
    Capsule splitting
    NA
    Fruit (pyxis) dehiscence
    NA
    Fruit features (Brassicaceae)
    NA
    Fruit length
    2.4–2.8 mm
    Fruit length relative to sepals
    the fruit is shorter than its associated sepals
    Fruit locules
    four
    Fruit shape
    the fruit is obloid (longer than wide and with rounded ends)
    Fruit stalk orientation
    NA
    Fruit type (general)
    the fruit is dry and splits open when ripe
    Fruit type (specific)
    the fruit is a schizocarp (when dry it splits into sections, each holding one or more seeds)
    Legumes (Fabaceae)
    NA
    Mericarp length
    2.4–2.8 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Ovary stipe length
    0 mm
    Placenta arrangement
    marginal
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    the fruit is not flattened
    Schizocarpic fruit segments
    4
    Seed number
    4
    Seed relative length
    the seed is longer than it is wide
    Seeds comose
    no hairs
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    the plant has a rhizome (a horizontal underground stem with roots growing from it)

  • Leaves

    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bracts
    the flower has one or more bracts associated with it
    Hairs on leaf stalk
    the petiole has hairs on it
    Hairs on underside of leaf
    • the underside of the leaf is fuzzy or hairy
    • the underside of the leaf is not hairy, or it has very few hairs
    Hairs on upper side of leaf
    • the upper side of the leaf is fuzzy or hairy
    • the upper side of the leaf is not hairy, or it has very few hairs
    Hooked hairs on underside of leaf
    no
    Inflated hairs on leaf
    the leaf blade does not have inflated hairs on it
    Leaf arrangement
    opposite: there are two leaves per node along the stem
    Leaf blade base
    the leaf has a distinct leaf stalk (petiole)
    Leaf blade base shape
    the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
    Leaf blade base symmetry
    the leaf blade base is symmetrical
    Leaf blade bloom
    the underside of the leaf has no noticeable bloom
    Leaf blade edges
    the edge of the leaf blade has teeth
    Leaf blade flatness
    the leaf is flat (planar) at the edges
    Leaf blade hairs
    • NA
    • the leaf blade has simple hairs with no glands, and not tangled or wooly
    Leaf blade length
    30–100 mm
    Leaf blade primary vein pattern
    the secondary veins branch off at intervals from the primary vein
    Leaf blade shape
    • the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
    • the leaf blade is linear (very narrow with more or less parallel sides)
    • the leaf blade is oblong (rectangular but with rounded ends)
    • the leaf blade is spatulate (spoon-shaped; narrow near the base, then suddenly widening to a rounded tip)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade texture
    the leaf blade is herbaceous (has a leafy texture)
    Leaf blade vein pattern
    the major veins of the leaf blade branch, but do not rejoin
    Leaf blade veins
    the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)
    Leaf blade width
    3–15 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf hair orientation
    • NA
    • the hairs are flat against the leaf surface, mostly pointing towards the leaf tip
    Leaf sheath length
    0 mm
    Leaf shiny
    the upper side of the leaf is dull or slightly shiny
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have leaf stalks
    Leaf stalk attachment to leaf
    the petiole attaches at the basal margin of the leaf blade
    Leaf stalk base
    the petiole base is narrow where it attaches to the stem
    Leaf teeth and lobes
    the leaf blade margin has forward-pointing teeth
    Leaf tip
    • the tip of the leaf blade is acute (sharply pointed)
    • the tip of the leaf blade is obtuse (bluntly pointed)
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf types
    • NA
    • There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaflet petiolules
    NA
    Leaves per node
    there are two leaves per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule edges
    NA
    Stipule features
    NA
    Stipule fused to leaf stalk
    NA
    Stipule length
    0 mm
    Stipule shape
    NA
    Stipules
    there are no stipules on the plant

  • Place

    Habitat
    terrestrial
    New England state
    • Connecticut
    • Massachusetts
    • Vermont
    Specific habitat
    • man-made or disturbed habitats
    • meadows or fields
    • ridges or ledges
    • sandplains or barrens

  • Scent

    Plant odor
    the plant does not have much of a smell

  • Stem, shoot, branch

    Branched tendrils
    NA
    Direction of stem hairs
    the hairs point mostly upwards to outwards
    Flowering stem cross-section
    the flowering stem is roughly square
    Hair between stem nodes
    the stem has hairs between the nodes
    Hairs between stem nodes
    the hairs on the stem are plain, without glands or branches, and not tangled
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    10–70 cm
    Stem bloom
    there is no powdery or waxy film on the stem
    Stem hair distribution
    the hairs on the stem are distributed more of less uniformly
    Stem nodes swollen
    the stem is not swollen at the nodes
    Stem orientation
    the stems are upright or angled outwards
    Stem roughness between nodes
    the stem does not feel rough
    Stem spacing
    the plant is solitary, or a few plants are growing together
    Stem succulence
    the stems are not succulent
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils
    Wings on stem
    the stem does not have wings on it

Wetland status

Not classified

In New England

Distribution

Connecticut
present
Maine
absent
Massachusetts
present
New Hampshire
absent
Rhode Island
absent
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

Connecticut
historical (S-rank: SH), special concern, extirpated (code: SC*)
Massachusetts
extremely rare (S-rank: S1), endangered (code: E)
Vermont
historical (S-rank: SH)

From Flora Novae Angliae dichotomous key

6.  Verbena simplex  Leym. NC

narrow-leaved vervain. Verbena angustifolia Michx. • CT, MA, VT. Dry fields, sandplains, ledges, quarries, road cuts. This species was reported from Grafton County, NH, by Magee and Ahles (1999). However, the herbarium specimens that voucher these occurrences (at least those formerly at HNH) are without locality information save for the pre-printed mention of Hanover, NH, on the Jesup label. It is doubtful that these collections originated in NH.

3×6. Verbena hastata × Verbena simplex → Verbena ×‌blanchardii Moldenke is a very rare vervain hybrid known from VT. It has 1–5 spikes at the summit of the stems or branches, moderately strigose stems (sparsely strigose in V. simplex), a corolla averaging larger than V. hastata (limb 3.8–5.3 mm wide), that is light blue or light purple (not deep blue as in V. hastata).

Native to North America?

Yes

Sometimes confused with

Verbena hispida:
leaf blades slightly tapered to the base, sometimes with an broad, ill-defined, petiole-like base, and spikes obloid to cylindric, mostly 3–7.5 times as long as wide (vs. V. simplex, with leaf blades conspicuously tapered and/or very narrow at the base, often with a short, but evident, petiole, and spikes cylindric to narrow-cylindric, mostly 7.5 or more times as long as wide).
Verbena stricta:
stems hirsute, often densely so, leaf blades elliptic to ovate or suborbicular, mostly 15–50 mm wide, and limb of corolla 7–9 mm wide (vs. V. simplex, with stems sparsely strigillose, leaf blades linear or narrow-oblong to lanceolate or narrow-spatulate, the primary ones 3–15 mm wide, and limb of corolla 4–6 mm wide).

Synonyms

  • Verbena angustifolia Michx.

Family

Verbenaceae

Genus

Verbena