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Pycnanthemum verticillatum — whorled mountain-mint

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Facts

Whorled mountain-mint is native to much of the eastern half of North America. There are two varieties, both of which occur in New England. This species can be difficult to separate from clustered mountain-mint (Pycnanthemum muticum) except on the basis of leaf blade shape. Since this character tends to be variable, examining multiple individuals may be helpful.

Habitat

Forests, meadows and fields

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
  • terrestrial
  • wetlands
New England state
  • Connecticut
  • Massachusetts
  • Rhode Island
  • Vermont
Flower petal color
white
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
opposite: there are two leaves per node along the stem
Leaf blade edges
  • the edge of the leaf blade has teeth
  • the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
Number of sepals, petals or tepals
  • there are five petals, sepals, or tepals in the flower
  • there are four petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
4
Fruit type (general)
the fruit is dry but does not split open when ripe
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla morphology
    the flower has two prominent lips
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower number
    At least 4
    Flower petal color
    white
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is branched above the base
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inflorescence width
    3–20 mm
    Inner tepals (Rumex)
    NA
    Marks on petals
    the petals have spots or streaks on them
    Nectar spur
    the flower has no nectar spurs
    Number of carpels
    2
    Number of pistils
    • 1
    • 4
    Number of sepals, petals or tepals
    • there are five petals, sepals, or tepals in the flower
    • there are four petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    white
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal number
    4–5
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appearance
    the sepals are green or brown, and leaf-like in texture
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal length
    4.3–5.2 mm
    Sepal number
    5
    Stamen attachment
    the stamens are attached at or near the bases of the petals or tepals
    Stamen number
    4
    Stamen relative length
    anything
    Stamens fused
    the stamens are not attached to one another
    Staminodes
    there are no staminodes on the flower
    Stigma position
    the stigmas are positioned at the tip of the style
    Style petal-like
    the styles are not petal-like
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    • the upper lip of the bilabiate corolla has one lobe
    • the upper lip of the bilabiate corolla has two lobes

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    NA
    Capsule splitting
    NA
    Fruit (pyxis) dehiscence
    NA
    Fruit beak length
    0 mm
    Fruit features (Brassicaceae)
    NA
    Fruit locules
    four
    Fruit shape
    the fruit is obloid (longer than wide and with rounded ends)
    Fruit type (general)
    the fruit is dry but does not split open when ripe
    Fruit type (specific)
    the fruit is a schizocarp (when dry it splits into sections, each holding one or more seeds)
    Legumes (Fabaceae)
    NA
    Mericarp length
    1–1.3 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    the fruit is not flattened
    Schizocarpic fruit segments
    4
    Seed number
    1–4
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    the plant has a rhizome (a horizontal underground stem with roots growing from it)

  • Leaves

    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bracts
    the flower has one or more bracts associated with it
    Hairs on underside of leaf
    the underside of the leaf is fuzzy or hairy
    Hairs on upper side of leaf
    • the upper side of the leaf is fuzzy or hairy
    • the upper side of the leaf is not hairy, or it has very few hairs
    Leaf arrangement
    opposite: there are two leaves per node along the stem
    Leaf blade base
    • the leaf has a distinct leaf stalk (petiole)
    • the leaf has no stalk
    Leaf blade base shape
    the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
    Leaf blade edges
    • the edge of the leaf blade has teeth
    • the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade length
    10–95 mm
    Leaf blade shape
    • the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
    • the leaf blade is linear (very narrow with more or less parallel sides)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade width
    3–15 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    • the leaves have leaf stalks
    • the leaves have no leaf stalks, but attach directly to the stem
    Leaf stalk length
    0–3 mm
    Leaf teeth and lobes
    • the edge of the leaf blade is entire (has no teeth or lobes)
    • the leaf blade margin has forward-pointing teeth
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaves per node
    there are two leaves per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule features
    NA
    Stipules
    there are no stipules on the plant

  • Place

    Habitat
    • terrestrial
    • wetlands
    New England state
    • Connecticut
    • Massachusetts
    • Rhode Island
    • Vermont
    Specific habitat
    • forests
    • meadows or fields

  • Scent

    Plant odor
    the plant has a pleasant smell, for example anise, fruit, mint or resin

  • Stem, shoot, branch

    Branched tendrils
    NA
    Flowering stem cross-section
    the flowering stem is roughly square
    Hair between stem nodes
    the stem has hairs between the nodes
    Hairs between stem nodes
    the hairs on the stem are plain, without glands or branches, and not tangled
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    40–120 cm
    Stem hair distribution
    the hairs on the stem are distributed more of less uniformly
    Stem orientation
    the stems are upright or angled outwards
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils

Wetland status

Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)

In New England

Distribution

Connecticut
present
Maine
absent
Massachusetts
present
New Hampshire
absent
Rhode Island
present
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

var. pilosum

Massachusetts
not applicable (S-rank: SNA)

var. verticillatum

Massachusetts
fairly widespread (uncertain) (S-rank: S4?)
Vermont
rare to uncommon (S-rank: S2S3)

Subspecies and varieties

Variety pilosum is known from CT, MA. Variety verticillatum is known from CT, MA, RI, VT; also reported from ME by Magee and Ahles (1999), but specimens are unknown.

From Flora Novae Angliae dichotomous key

5.  Pycnanthemum verticillatum (Michx.) Pers. N

whorled mountain-mint.  5a. Koellia pilosa (Nutt.) Britt.; Pycnanthemum pilosum Nutt.;  
5b. Koellia verticillata (Michx.) Kuntze; Pycnanthemum virginianum (L.) T. Dur. & B.D. Jackson ex B.L. Robins. & Fern. var. verticillatum (Michx.) Boivin • CT, MA, RI, VT; also reported from ME by Magee and Ahles (1999), but specimens are unknown. Mesic to wet-mesic forests, fields, meadows, and roadsides.

1a.  Faces of upper stem internodes conspicuously pilose with spreading hairs; pubescence of abaxial leaf blade conspicuous and uniform across the surface 
 … 5a. P. verticillatum var. pilosum (Nutt.) Cooperrider

1b.  Faces of upper stem internodes moderately to sparsely pubescent with short, pilose hairs; pubescence of abaxial leaf blade chiefly or entirely confined to the veins 
 … 5b. P. verticillatum var. verticillatum

Variety pilosum is known from CT, MA. Variety verticillatum is known from CT, MA, RI, VT; 
also reported from ME by Magee and Ahles (1999), but specimens are unknown.

Native to North America?

Yes and no (some introduced)

Sometimes confused with

Pycnanthemum torrei:
bracts and leaf blades glabrous on the adaxial surface or sometimes with minute, stiff scabrules on the adaxial bract surface (vs. P. verticillatum, with the bracts, and often also the leaf blades, pubescent on the adaxial surface with minute, soft hairs that are best viewed at 20x).
Pycnanthemum virginianum:
bracts and leaf blades glabrous on the adaxial surface or sometimes with minute, stiff scabrules on the adaxial bract surface, and stems pubescent primarily on the angles (vs. P. verticillatum, with the bracts, and often also the leaf blades, pubescent on the adaxial surface with minute, soft hairs that are best viewed at 20x, and stems pubescent on the faces as well as the angles).

Synonyms

  • Koellia verticillata (Michx.) Kuntze
  • Pycnanthemum virginianum (L.) T. Dur. & B.D. Jackson ex B.L. Robins. & Fern. var. verticillatum (Michx.) Boivin

Family

Lamiaceae

Genus

Pycnanthemum