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Native Plant Trust: Go Botany Discover thousands of New England plants

Linnaea borealis — American twinflower

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Facts

Research has shown that American twinflower cannot pollinate close relatives, and needs less closely-related individuals nearby in order to successfully set seed. Thus, if a population becomes small or isolated from others, it is more likely to die out.

Habitat

Alpine or subalpine zones, forests, talus and rocky slopes, woodlands

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
  • terrestrial
  • wetlands
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Flower petal color
  • pink to red
  • white
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
opposite: there are two leaves per node along the stem
Leaf blade edges
the edge of the leaf blade has teeth
Flower symmetry
there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
Number of sepals, petals or tepals
there are five petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
4
Fruit type (general)
the fruit is dry but does not split open when ripe
Fruit length
3 mm
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Anther spurs
    the anthers do not have spurs on them
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Flower description
    the flower has an inferior ovary, with or without a hypanthium
    Flower length
    10–15 mm
    Flower number
    2
    Flower orientation
    the flower bends downwards or hangs downwards
    Flower petal color
    • pink to red
    • white
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is knob-like at the tip, and unbranched
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Hairs on flower stalk
    the flower stalk has hairs on it
    Hairs on inflorescence
    at least some of the hairs on the axis of the inflorescence have glands
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inner tepals (Rumex)
    NA
    Length of flower stalk
    10–20 mm
    Length of peduncle
    45–80 mm
    Marks on petals
    the petals have spots or streaks on them
    Nectar spur
    the flower has no nectar spurs
    Number of branches in umbel
    0
    Number of sepals, petals or tepals
    there are five petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is below the point of petal and/or sepal attachment
    Perianth shape
    the perianth is campanulate (bell-shaped, with a tube about as long as wide, flaring at the mouth)
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    • pink to red
    • white
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal hairs on inner/upper surface
    there are hairs on the inner/upper petal surface
    Petal length
    10–15 mm
    Petal number
    5
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appearance
    the sepals are green or brown, and leaf-like in texture
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal color
    pink to red
    Sepal number
    5
    Sepal shape
    the sepal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)
    Sepal uniformity
    all the sepals are about the same size
    Stamen morphology
    the stamens within a cycle differ in length or width
    Stamen number
    4
    Stamen relative length
    anything
    Staminodes
    there are no staminodes on the flower
    Style petal-like
    the styles are not petal-like
    Style relative length
    the stigma protrudes beyond the mouth of the corolla
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    NA

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule splitting
    the capsule splits by three main valves, teeth or pores
    Carpel beak length
    0 mm
    Fruit (pyxis) dehiscence
    NA
    Fruit beak length
    0 mm
    Fruit features (Brassicaceae)
    NA
    Fruit length
    3 mm
    Fruit locules
    three
    Fruit type (general)
    the fruit is dry but does not split open when ripe
    Fruit type (specific)
    the fruit is a nut (dry and indehiscent, with a hard wall, usually containing only one seed and usually subtended by an involucre)
    Hair type on fruit
    the hairs on the fruits have glands at their tips
    Hairs on fruit
    the fruits have hairs on them
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Seed number
    1
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    • the plant is a subshrub (small shrub with partially herbaceous stems)
    • the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    • the plant does not have stolons
    • the plant has stolons
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines

  • Leaves

    Bracteole length
    1 mm
    Bracteole number (Apiaceae)
    0
    Bracteole shape
    the bracteoles are lanceolate (widest below the middle, and tapering at both ends)
    Bracteoles
    the plant has bracteoles between the primary bracts and the flowers
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bract texture
    the floral bracts are green, with a leaf-like texture
    Floral bracts
    the flower has one or more bracts associated with it
    Flower bract length
    1.5–2 mm
    Hooked hairs on underside of leaf
    no
    Inflated hairs on leaf
    the leaf blade does not have inflated hairs on it
    Leaf arrangement
    opposite: there are two leaves per node along the stem
    Leaf blade base
    the leaf has a distinct leaf stalk (petiole)
    Leaf blade base shape
    the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
    Leaf blade base symmetry
    the leaf blade base is symmetrical
    Leaf blade bloom
    the underside of the leaf has no noticeable bloom
    Leaf blade edges
    the edge of the leaf blade has teeth
    Leaf blade flatness
    the leaf is flat (planar) at the edges
    Leaf blade length
    5–16 mm
    Leaf blade primary vein pattern
    the secondary veins branch off at intervals from the primary vein
    Leaf blade shape
    • the leaf blade is orbicular (roughly circular, as wide as long)
    • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade texture
    the leaf blade is coriaceous (has a firm, leathery texture)
    Leaf blade vein pattern
    the major veins of the leaf blade branch, but do not rejoin
    Leaf blade veins
    the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)
    Leaf blade width
    4–10 mm
    Leaf duration
    the leaves remain green all winter
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf shiny
    the upper side of the leaf is dull or slightly shiny
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have leaf stalks
    Leaf stalk attachment to leaf
    the petiole attaches at the basal margin of the leaf blade
    Leaf stalk length
    2–3 mm
    Leaf teeth and lobes
    • the leaf blade margin has outward-pointing teeth
    • the leaf blade margin has rounded teeth
    Leaf tip
    the tip of the leaf blade is obtuse (bluntly pointed)
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf types
    NA
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaflet petiolules
    NA
    Leaves per node
    there are two leaves per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule edges
    NA
    Stipule features
    NA
    Stipule fused to leaf stalk
    NA
    Stipule length
    0 mm
    Stipule shape
    NA
    Stipules
    there are no stipules on the plant

  • Place

    Habitat
    • terrestrial
    • wetlands
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • alpine or subalpine zones
    • forests
    • talus or rocky slopes
    • woodlands

  • Scent

    Plant odor
    the plant does not have much of a smell

  • Stem, shoot, branch

    Branched tendrils
    NA
    Flowering stem cross-section
    the flowering stem is circular, or with lots of small angles so that it is roughly circular
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Stem bloom
    there is no powdery or waxy film on the stem
    Stem nodes swollen
    the stem is not swollen at the nodes
    Stem orientation
    the stems trail at the base, but may turn upwards at the tips
    Stem succulence
    the stems are not succulent
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils
    Wings on stem
    the stem does not have wings on it

Wetland status

Occurs in wetlands or non-wetlands. (Wetland indicator code: FAC)

In New England

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
present
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

Connecticut
unranked (S-rank: SNR)
Maine
unranked (S-rank: SNR)

ssp. americana

Connecticut
extremely rare (S-rank: S1), endangered (code: E)
Massachusetts
uncommon (S-rank: S3)
Rhode Island
historical (S-rank: SH), state historical (code: SH)

ssp. longiflora

Maine
sir (S-rank: sir)

Subspecies and varieties

Our subspecies is Linnaea borealis L. ssp. americana (Forbes) Hultén ex Clausen.

From Flora Novae Angliae dichotomous key

1.  Linnaea borealis L. ssp. americana (Forbes) Hultén ex Clausen N

American twinflower. Linnaea americana Forbes; L. borealis L. var. americana (Forbes) Rehd. • CT, MA, ME, NH, RI, VT; becoming rare in southern New England. Mesic to wet-mesic, evergreen and mixed evergreen-deciduous forests, ascending to boreal situations.

Native to North America?

Yes

Synonyms

  • Linnaea americana Forbes
  • Linnaea borealis var. americana (Forbes) Rehd.

Family

Caprifoliaceae

Genus

Linnaea