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Linaria dalmatica — Dalmatian toadflax

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Facts

Dalmatian toadflax, a widespread weed in North America, was introduced from the steppes of its native Eurasia in the late 1800s. It is a short-lived, perennial herb with yellow flowers and waxy, vaguley heart-shaped leaves arrayed on stems up to 90 cm (3 feet) tall. Its deep taproot and extensive lateral rooting system allow it to capture water efficiently, resprout after fire, and withstand the attempts of grazers (or gardeners) to pull it up; it can also reproduce vegetatively from buds off the roots.

Habitat

Anthropogenic (man-made or disturbed habitats), meadows and fields

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
terrestrial
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Flower petal color
yellow
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
  • alternate: there is one leaf per node along the stem
  • opposite: there are two leaves per node along the stem
  • whorled: there are three or more leaves per node along the stem
Leaf blade edges
  • the edge of the leaf blade has teeth
  • the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
Number of sepals, petals or tepals
there are five petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
4
Fruit type (general)
the fruit is dry and splits open when ripe
Fruit length
3–7 mm
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Anther spurs
    the anthers do not have spurs on them
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla morphology
    the flower has two prominent lips
    Corolla palate
    yes
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Filament surface
    the filament is smooth, with no hairs or scales
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower petal color
    yellow
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is knob-like at the tip, and unbranched
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Inner tepals (Rumex)
    NA
    Nectar spur
    the flower has one or more nectar spurs
    Number of carpels
    2
    Number of pistils
    • 1
    • 2
    Number of sepals, petals or tepals
    there are five petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    yellow
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal number
    5
    Petal tips (Cuscuta)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal number
    5
    Sepals fused only to sepals
    the sepals are fused to each other (not other flower parts), at least near their bases
    Stamen attachment
    the stamens are attached at or near the bases of the petals or tepals
    Stamen number
    4
    Stamen position relative to petals
    the stamens are lined up with the sepals
    Stigma position
    the stigmas are positioned at the tip of the style
    Style petal-like
    the styles are not petal-like
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    the upper lip of the bilabiate corolla has two lobes

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    the capsule has no prominent ribs or wings
    Capsule splitting
    the capsule splits by two main valves, teeth or pores
    Fruit (pyxis) dehiscence
    NA
    Fruit features (Brassicaceae)
    NA
    Fruit length
    3–7 mm
    Fruit locules
    two
    Fruit type (general)
    the fruit is dry and splits open when ripe
    Fruit type (specific)
    the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines

  • Leaves

    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Hairs on underside of leaf
    the underside of the leaf is not hairy, or it has very few hairs
    Hairs on upper side of leaf
    the upper side of the leaf is not hairy, or it has very few hairs
    Leaf arrangement
    • alternate: there is one leaf per node along the stem
    • opposite: there are two leaves per node along the stem
    • whorled: there are three or more leaves per node along the stem
    Leaf blade base
    the leaf has no stalk
    Leaf blade base shape
    the base of the leaf blade is rounded
    Leaf blade edges
    • the edge of the leaf blade has teeth
    • the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade length
    20–50 mm
    Leaf blade shape
    • the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
    • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have no leaf stalks, but attach directly to the stem
    Leaf teeth and lobes
    • the edge of the leaf blade is entire (has no teeth or lobes)
    • the leaf blade margin has outward-pointing teeth
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaves per node
    • there are four leaves per node along the stem
    • there are three leaves per node along the stem
    • there are two leaves per node along the stem
    • there is one leaf per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule features
    NA
    Stipules
    there are no stipules on the plant

  • Place

    Habitat
    terrestrial
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • man-made or disturbed habitats
    • meadows or fields

  • Scent

    Plant odor
    the plant does not have much of a smell

  • Stem, shoot, branch

    Branched tendrils
    NA
    Flowering stem cross-section
    the flowering stem is circular, or with lots of small angles so that it is roughly circular
    Hair between stem nodes
    the stem has no hairs between the nodes
    Hairs between stem nodes
    the stem has no hairs between the nodes
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    40–120 cm
    Stem bloom
    the stem has a powdery or waxy film on it that can be rubbed away
    Stem roughness between nodes
    the stem does not feel rough
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils

Wetland status

Not classified

In New England

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
absent
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

ssp. dalmatica

Massachusetts
not applicable (S-rank: SNA)
Vermont
not applicable (S-rank: SNA)

Subspecies and varieties

Linaria dalmatica ssp. dalmatica is known from CT, MA, ME, NH, RI, VT. Linaria dalmatica ssp. macedonica (Griseb.) D.A. Sutton is known from NH.

From Flora Novae Angliae dichotomous key

1.  Linaria dalmatica (L.) P. Mill. E

Dalmatian toadflax. 1a. Antirrhinum dalmaticum L.; L. genistifolia (L.) P. Mill. ssp. dalmatica (L.) Maire & Petitm.; 1b. L. macedonica Griseb.; L. dalmatica var. macedonica (Griseb.) Vandas • CT, MA, ME, NH, RI, VT. Fields, roadsides, waste areas.

1a. Pedicels 3–8 mm long in flower, 4–10(–13) mm long in fruit; floral bracts usually +/- equal to or longer than the fruiting pedicels in length; corollas 25–38(–42) mm long, with nectary spurs 11–24 mm long that vary in length from shorter than to longer than the rest of the corolla ..... 1a. L. dalmatica ssp. dalmatica

1b. Pedicels 7–22 mm long in flower, (7–)12–32 mm long in fruit; floral bracts usually shorter than the fruiting pedicels; corollas 30–49 mm long, with nectary spurs 19–30 mm long that vary in length from +/- equal to or longer than the rest of the corolla ..... 1b. L. dalmatica ssp. macedonica (Griseb.) D.A. Sutt.

Subspecies dalmatica is known from CT, MA, ME, NH, RI, VT. Subspecies macedonica is known from NH.

Native to North America?

No

Sometimes confused with

Linaria genistifolia:
corolla 13–22 mm long and leaf blades 3–15 mm wide, narrow-lanceolate to broad-lanceolate, mostly 6–12 times as long as wide (vs. L. dalmatica, with the corolla 20–50 mm long and leaf blades 10–40 mm wide, broad-lanceolate to ovate, mostly 2–4 times as long as wide).

Synonyms

  • Antirrhinum dalmaticum L.
  • Linaria genistifolia (L.) P. Mill. ssp. dalmatica (L.) Maire & Petitm.

Family

Plantaginaceae

Genus

Linaria