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- Gentianaceae
- Gentiana
- Gentiana clausa
Gentiana clausa — closed gentian, meadow bottle gentian
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Facts
Gentians are named after King Gentius of Illyria who is credited with discovering medicinal properties of the root. The flowers do not open and are pollinated by insects that are strong enough to force their way in.
Habitat
Anthropogenic (man-made or disturbed habitats), floodplain (river or stream floodplains), forests, meadows and fields
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
Found this plant? Take a photo and post a sighting.
Characteristics
- Habitat
-
- terrestrial
- wetlands
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Vermont
- Flower petal color
-
- blue to purple
- white
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf arrangement
- opposite: there are two leaves per node along the stem
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Number of sepals, petals or tepals
-
- there are five petals, sepals, or tepals in the flower
- there are four petals, sepals, or tepals in the flower
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Stamen number
-
- 4
- 5
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit length
- At least 10 mm
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Anther attachment
- the anther is attached by its base to the filament
- Anther opening
- the anthers have narrow slits or furrows that run lengthwise along the anthers
- Anther spurs
- the anthers do not have spurs on them
- Anther tube length
- 0 mm
- Calyx growth after flowering
- the calyx does not grow to cover or partially cover the fruit
- Calyx symmetry
- there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
- Carpels fused
- the carpels are fused to one another
- Cilia on petals
- the petal margins do not have cilia
- Cleistogamous flowers
- there are no cleistogamous flowers on the plan
- Corolla palate
- no
- Corona lobe length
- 0 mm
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Filament surface
- the filament is smooth, with no hairs or scales
- Flower description
- the flower has a superior ovary, and lacks a hypanthium
- Flower length
- 5–55 mm
- Flower number
- At least 1
- Flower orientation
- the flower points upwards or is angled outwards
- Flower petal color
-
- blue to purple
- white
- Flower reproductive parts
- the flower has both pollen- and seed-producing parts
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Flowers sunken into stem
- no
- Form of style
- the style is lobed at the tip, and unbranched
- Fringed petal edges
- the petals are not fringed
- Fused stamen clusters
- NA
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Hairs on flower stalk
- the flower stalk has no hairs on it
- Hairs on inflorescence
- the axis of the inflorescence has no hairs on it
- Horns in hoods (Asclepias)
- NA
- Hypanthium
- the flower does not have a hypanthium
- Hypanthium length
- 0 mm
- Inflorescence one-sided
- the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
- Inner tepals (Rumex)
- NA
- Length of flower stalk
- At least 0 mm
- Length of peduncle
- 0 mm
- Marks on petals
- there are no noticeable marks on the petals
- Nectar spur
- the flower has no nectar spurs
- Number of branches in umbel
- 0
- Number of carpels
- 2
- Number of pistils
- 1
- Number of sepals, petals or tepals
-
- there are five petals, sepals, or tepals in the flower
- there are four petals, sepals, or tepals in the flower
- Number of styles
- 0–1
- Ovary position
- the ovary is above the point of petal and/or sepal attachment
- Perianth shape
-
- the perianth is campanulate (bell-shaped, with a tube about as long as wide, flaring at the mouth)
- the perianth is cylindric (tube-shaped)
- the perianth is funnelform (funnel-shaped)
- Petal and sepal arrangement
- the flower includes two cycles of petal- or sepal-like structures
- Petal and sepal colors
-
- blue to purple
- white
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal base
- NA
- Petal folding in bud
- the petals in bud are rolled like an umbrella, each petal having one edge tucked inside and the other edge exposed (convolute)
- Petal folds or pleats
- the petals of the flower have folds or plaits on them
- Petal glandular dots or scales
- no
- Petal hairs (Viola)
- NA
- Petal hairs on inner/upper surface
- there are no hairs on the inner/upper petal surface
- Petal length relative to sepals
- the petals are longer than the sepals
- Petal nectaries
- the petals have at least one nectary
- Petal number
- Up to 5
- Petal tip shape
- the petal tip is rounded
- Petal tips (Cuscuta)
- NA
- Petal width
- At least 3 mm
- Raceme attachment (Veronica)
- NA
- Reproductive system
- all the flowers have both carpels and stamens (synoecious)
- Scales inside corolla
- no
- Sepal and petal color
- the sepals are different from the petals
- Sepal appearance
- the sepals are green or brown, and leaf-like in texture
- Sepal appendages
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Sepal auricles
- the sepals have no auricles
- Sepal cilia
- the sepals have cilia on their edges
- Sepal color
- green to brown
- Sepal number
- 4–5
- Sepal shape
-
- the sepal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)
- the sepal outline is obovate (egg-shaped, but with the widest point above the middle of the leaf blade)
- Sepal texture
- the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
- Sepal uniformity
- all the sepals are about the same size
- Sepals fused only to sepals
- the sepals are fused to each other (not other flower parts), at least near their bases
- Spur length
- 0 mm
- Spur number
- NA
- Stamen appendages
- stamen appendages are absent
- Stamen attachment
- the stamens are attached at or near the bases of the petals or tepals
- Stamen lengths differ
- the stamens are all approximately the same length
- Stamen morphology
- the stamens within each cycle are the same
- Stamen number
-
- 4
- 5
- Stamen position relative to petals
- the stamens are lined up with the petals
- Stamen relative length
- anything
- Stamens fused
- the stamens are not attached to one another
- Staminodes
- there are no staminodes on the flower
- Stigma position
- the stigmas are positioned at the tip of the style
- Style petal-like
- the styles are not petal-like
- Style relative length
- the stigma does not protrude beyond the mouth of the corolla
- Surface of ovary
- the ovary surface is textured with tiny points, bumps or wrinkles
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene relative orientation
- NA
- Achene shape
- NA
- Achene surface (Polygonum)
- NA
- Achene type
- NA
- Berry color
- NA
- Capsule color (Viola)
- NA
- Capsule ribs
- the capsule has two prominent ribs or wings
- Capsule splitting
- the capsule splits by two main valves, teeth or pores
- Carpel beak length
- 0 mm
- Fruit (pyxis) dehiscence
- NA
- Fruit features (Brassicaceae)
- NA
- Fruit length
- At least 10 mm
- Fruit locules
- one
- Fruit shape
-
- the fruit is ellipsoid (widest in the middle and tapering to each end)
- the fruit is obovoid (egg-shaped, but with the widest point above the middle)
- the fruit is roughly cylindrical (with parallel sides that do not taper, and flat across the top and bottom)
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit type (specific)
- the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)
- Legumes (Fabaceae)
- NA
- Mericarp length
- 0 mm
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
- NA
- Ovary stipe
- the ovary or fruit has a stipe
- Placenta arrangement
- the plant has parietal placentation, where ovules develop on the wall or slight outgrowths of the wall forming broken partitions within a compound ovary
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- NA
- Schizocarpic fruit segments
- 0
- Seed length
- 0.6–2.2 mm
- Seed surface
-
- the seed has a surface characteristic other than those listed
- the seed is covered with reticulate markings (a netlike pattern due to splitting and rejoining of lines or ridges)
- Seeds comose
- no hairs
- Septum in fruit (Brassicaceae)
- NA
- Wings on fruit
- the fruit does not have wings on it
- prickles on fruits
- the fruits do not have thorn-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades do not have glandular dots or scales
- Sap
- the sap is clear and watery
- Sap color
- the sap is clear
-
Growth form
- Growth form
- the plant is an herb (it has self-supporting stems)
- Lifespan
- the plant lives more than two years
- Parasitism
- the plant is not parasitic
- Plant color
- the leaves or young stems of the plant are green
- Plants darken when dry
- no
- Spines on plant
- the plant has no spines
- Underground organs
-
- there are only slender roots on the plant
- there is a thickened taproot on the plant
-
Leaves
- Bracteole number (Apiaceae)
- 0
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length (compound lvs only)
- 0 mm
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Final leaf segment width (compound lvs only)
- 0 mm
- Hairs on underside of leaf
- the underside of the leaf is not hairy, or it has very few hairs
- Hairs on upper side of leaf
- the upper side of the leaf is not hairy, or it has very few hairs
- Hooked hairs on underside of leaf
- no
- Inflated hairs on leaf
- the leaf blade does not have inflated hairs on it
- Leaf arrangement
- opposite: there are two leaves per node along the stem
- Leaf blade base
-
- the leaf has a distinct leaf stalk (petiole)
- the leaf has no stalk
- Leaf blade base shape
- the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
- Leaf blade base symmetry
- the leaf blade base is symmetrical
- Leaf blade bloom
- the underside of the leaf has no noticeable bloom
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf blade flatness
- the leaf is flat (planar) at the edges
- Leaf blade hairs
- the leaf blade has simple hairs with no glands, and not tangled or wooly
- Leaf blade length
- 5–150 mm
- Leaf blade primary vein pattern
- the secondary veins branch off at intervals from the primary vein
- Leaf blade shape
-
- the leaf blade is linear (very narrow with more or less parallel sides)
- the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf blade texture
- the leaf blade is herbaceous (has a leafy texture)
- Leaf blade vein pattern
- the major veins of the leaf blade branch, but do not rejoin
- Leaf blade width
- Up to 42 mm
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are green, with an expanded blade and a leaf-like texture
- Leaf hair orientation
- NA
- Leaf shiny
- the upper side of the leaf is dull or slightly shiny
- Leaf spines
- there are no spines on the leaf edges
- Leaf stalk
-
- the leaves have leaf stalks
- the leaves have no leaf stalks, but attach directly to the stem
- Leaf stalk attachment to leaf
-
- NA
- the petiole attaches at the basal margin of the leaf blade
- Leaf teeth and lobes
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf tip
- the tip of the leaf blade is acuminate (tapers to a long, thin point)
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf types
- There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).
- Leaf variation
- the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
- there are two leaves per node along the stem
- Pinnately compound leaf type
- NA
- Specific leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Stipels
- NA
- Stipule edges
- NA
- Stipule features
- NA
- Stipule fused to leaf stalk
- NA
- Stipule length
- 0 mm
- Stipule shape
- NA
- Stipules
- there are no stipules on the plant
- Teeth per side of leaf blade
- 0
-
Place
- Habitat
-
- terrestrial
- wetlands
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Vermont
- Specific habitat
-
- forests
- man-made or disturbed habitats
- meadows or fields
- river or stream floodplains
-
Scent
- Plant odor
- the plant does not have much of a smell
-
Stem, shoot, branch
- Branched tendrils
- NA
- Direction of stem hairs
- NA
- Flowering stem cross-section
-
- the flowering stem is circular, or with lots of small angles so that it is roughly circular
- the flowering stem is roughly square
- Hair between stem nodes
- the stem has no hairs between the nodes
- Hairs between stem nodes
- the stem has no hairs between the nodes
- Hooked hairs on stem between nodes
- no
- Leaves on stem
- there is at least one full leaf above the base of the flowering stem
- Length of hairs between stem nodes
- 0 mm
- Stem hair distribution
- NA
- Stem orientation
- the stems are upright or angled outwards
- Stem roughness between nodes
- the stem does not feel rough
- Stem spacing
- the plant is solitary, or a few plants are growing together
- Stem succulence
- the stems are not succulent
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
- Wings on stem
- the stem does not have wings on it
Wetland status
Usually occurs in wetlands, but occasionally in non-wetlands. (Wetland indicator code: FACW)
In New England
Distribution
- Connecticut
- present
- Maine
- present
- Massachusetts
- present
- New Hampshire
- present
- Rhode Island
- present
- Vermont
- present
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Massachusetts
- fairly widespread (S-rank: S4)
- Rhode Island
- concern (code: C)
From Flora Novae Angliae dichotomous key
2. Gentiana clausa Raf. N
meadow bottle gentian. CT, MA, ME, NH, RI, VT. Meadows, mesic forests, clearings, riparian forests, roadside banks.
Native to North America?
Yes
Sometimes confused with
- Gentiana andrewsii:
- appendages in the sinuses of the corolla fimbriate, longer than the corolla lobes, and lobes of the corolla truncate and often apiculate, narrower than the plaits (vs. G. clausa, with appendages in the sinuses of the corolla 2- or 3-cleft, about as long as the corolla lobes, and lobes of the corolla broad-rounded at the apex, about as wide as the plaits).
- Gentiana linearis:
- margins of leaf blades lacking marginal cilia or with very minute, scattered cilia and leaf blades narrow-lanceolate (vs. G. clausa, with the margins of leaf blades with abundant cilia that are visible with low magnification and leaf blades lanceolate to narrow-ovate).