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- Dendrolycopodium dendroideum
Dendrolycopodium dendroideum — prickly tree-clubmoss
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Facts
The branched stem of prickly tree-clubmoss give it the appearance of a miniature tree, and the leaves on the lower part of the upright stem are, in fact, prickly.
Habitat
Forest edges, forests, woodlands
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
Found this plant? Take a photo and post a sighting.
Characteristics
- Habitat
- terrestrial
- New England state
- 
                                
                                    - Connecticut
- Maine
- Massachusetts
- New Hampshire
- Vermont
 
- Leaf shape
- the vegetative leaves are short and scale-like
- Spore leaf arrangement
- the sporophylls are located on spore cones at the tips of the shoots or branches
- Form of shoot
- the plant has an upright stem, with branches, those branches having further branches, and so on (tree-like)
- Horizontal stem
- the horizontal stem is under ground
- Leaf differences
- the vegetative leaves within a node are all similar in size and shape
- Teeth on leaf edges
- the edges of the vegetative leaves have no teeth
- Constriction zones
- 
                                
                                    - there are constricted zones on the vertical stem where the leaves are smaller smaller or closer together
- there are no constrictions on the horizontal stem with smaller leaves
 
- Spore leaf length
- 3.1–4.1 mm
- Leaf outline
- the vegetative leaves are long and very narrow (linear)
- 
                        Clonal plantlets- Gemma arrangement
- NA
 - Gemma shape
- NA
 - Gemma width
- 0 mm
 
- 
                        Leaves- Leaf differences
- the vegetative leaves within a node are all similar in size and shape
 - Leaf length
- 2–4 mm
 - Leaf orientation
- 
                                
                                    - the vegetative leaves spread away from the stem
- the vegetative leaves spread slightly away from the stem, at a steep angle
 
 - Leaf outline
- the vegetative leaves are long and very narrow (linear)
 - Leaf ranks
- 6 or 7
 - Leaf shape
- the vegetative leaves are short and scale-like
 - Spore leaf length
- 3.1–4.1 mm
 - Teeth on leaf edges
- the edges of the vegetative leaves have no teeth
 
- 
                        Place- Habitat
- terrestrial
 - New England state
- 
                                
                                    - Connecticut
- Maine
- Massachusetts
- New Hampshire
- Vermont
 
 - Specific habitat
- 
                                
                                    - edges of forests
- forests
- woodlands
 
 
- 
                        Spores or spore cones- Cone base at stem
- the base of the spore-cone has a distinct stalk
 - Cone stalk branching
- NA
 - Cone thickness
- 0 mm
 - Length of cone
- 12–55 mm
 - Number of cones
- 1–14
 - Quillwort itssue covering spores
- NA
 - Same or different spores
- there is only one type of spore present
 - Spore girdle
- there is no girdle alongside the equatorial ridge
 - Spore leaf arrangement
- the sporophylls are located on spore cones at the tips of the shoots or branches
 - Spore leaf lifespan
- the sporophylls wither and fall off at the end of the growing season
 - Spore leaf orientation
- the sporophylls are pressed against the spore cone
 - Spore leaf shape
- the spore-bearing leaves are small and scale-like
 - Spore leaf teeth
- The edges of the spore-bearing leaves are smooth, and without teeth
 - Spore texture
- the spore surface has a net-like pattern on it (reticulate)
 - Sterile tip of cone
- the spore cone does not have a slender, sterile tip (the whole cone produces spores)
 
- 
                        Stem, shoot, branch- Branch cross-section
- the outermost level of branches are round, elliptic or semicircular in cross-section
 - Branch form
- the branches are smaller than the main stem
 - Constriction zones
- 
                                
                                    - there are constricted zones on the vertical stem where the leaves are smaller smaller or closer together
- there are no constrictions on the horizontal stem with smaller leaves
 
 - Form of shoot
- the plant has an upright stem, with branches, those branches having further branches, and so on (tree-like)
 - Horizontal stem
- the horizontal stem is under ground
 - Stem height
- Up to 260 mm
 
Wetland status
Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)
In New England
Distribution
- Connecticut
- present
- Maine
- present
- Massachusetts
- present
- New Hampshire
- present
- Rhode Island
- present
- Vermont
- present
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Massachusetts
- fairly widespread (S-rank: S4)
From Flora Novae Angliae dichotomous key
1. Dendrolycopodium dendroideum (Michx.) A. Haines N
prickly tree-clubmoss. Lycopodium dendroideum Michx.; L. obscurum L. var. dendroideum (Michx.) D.C. Eat.; L. obscurum L. var. hybridum Farw. • CT, MA, ME, NH, VT; throughout most of the region except southeastern New England. Interior and edges of hardwood and mixed conifer-hardwood forests and woodlands.
1×2. Dendrolycopodium dendroideum × Dendrolycopodium hickeyi → This rare tree-clubmoss hybrid is known from CT, ME, VT.
1×3. Dendrolycopodium dendroideum × Dendrolycopodium obscurum → This rare tree-clubmoss hybrid is known from CT, MA, VT.
Native to North America?
Yes
Sometimes confused with
- Dendrolycopodium hickeyi:
- leaves of main axis ascending to appressed (vs. D. dendroideum, with widely ascending to spreading).
Synonyms
- Lycopodium dendroideum Michx.
- Lycopodium obscurum L. var. dendroideum (Michx.) D.C. Eat.
- Lycopodium obscurum L. var. hybridum Farw.
![Shoots: Dendrolycopodium dendroideum. ~ By Marilee Lovit. ~ Copyright © 2025 Marilee Lovit. ~ lovitm[at]gmail.com](https://newfs.s3.amazonaws.com/taxon-images-239x239/Lycopodiaceae/dendrolycopodium-dendroideum-sh-mlovit-c.jpg) 
                         
                        ![Branches: Dendrolycopodium dendroideum. ~ By Arthur Haines. ~ Copyright © 2025 Arthur Haines. ~ arthur.d.haines[at]gmail.com](https://newfs.s3.amazonaws.com/taxon-images-239x239/Lycopodiaceae/dendrolycopodium-dendroideum-br-ahaines-a.jpg) 
                        ![Spore cones: Dendrolycopodium dendroideum. ~ By Marilee Lovit. ~ Copyright © 2025 Marilee Lovit. ~ lovitm[at]gmail.com](https://newfs.s3.amazonaws.com/taxon-images-239x239/Lycopodiaceae/dendrolycopodium-dendroideum-sc-mlovit-b.jpg) 
                        ![Shoots: Dendrolycopodium dendroideum. ~ By Arthur Haines. ~ Copyright © 2025 Arthur Haines. ~ arthur.d.haines[at]gmail.com](https://newfs.s3.amazonaws.com/taxon-images-239x239/Lycopodiaceae/dendrolycopodium-dendroideum-sh-ahaines-b.jpg) 
                        ![Plant form: Dendrolycopodium dendroideum. ~ By Marilee Lovit. ~ Copyright © 2025 Marilee Lovit. ~ lovitm[at]gmail.com](https://newfs.s3.amazonaws.com/taxon-images-239x239/Lycopodiaceae/dendrolycopodium-dendroideum-ha-mlovit-a.jpg) 
                         
                        ![Vegetative leaves: Dendrolycopodium dendroideum. ~ By Arthur Haines. ~ Copyright © 2025 Arthur Haines. ~ arthur.d.haines[at]gmail.com](https://newfs.s3.amazonaws.com/taxon-images-239x239/Lycopodiaceae/dendrolycopodium-dendroideum-tr-ahaines-d.jpg) 
                        ![Vegetative leaves: Dendrolycopodium dendroideum. ~ By Elizabeth Farnsworth. ~ Copyright © 2025 New England Wild Flower Society. ~ Image Request, images[at]newenglandwild.org](https://newfs.s3.amazonaws.com/taxon-images-239x239/Lycopodiaceae/dendrolycopodium-dendroideum-tr-efarnsworth.jpg) 
                         
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