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- Bartonia iodandra
Bartonia iodandra — purple screwstem
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Facts
Purple screwstem is a rare northeastern endemic found only in New England and eastern Canada. It inhabits swamps, peatlands, meadows and abandoned borrow pits. It is very similar in appearance to the much more common twining screwstem (Bartonia paniculata).
Habitat
Anthropogenic (man-made or disturbed habitats), bogs, fens, meadows and fields, swamps
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
Found this plant? Take a photo and post a sighting.
Characteristics
- Habitat
-
- terrestrial
- wetlands
- Flower petal color
-
- pink to red
- white
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf arrangement
- alternate: there is one leaf per node along the stem
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Number of sepals, petals or tepals
- there are four petals, sepals, or tepals in the flower
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Stamen number
- 4
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit length
- 4–6 mm
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Anther color
- there is a noticeable pink, reddish or purplish tint to the anthers
- Anther length
- 0.4–1 mm
- Anther opening
- the anthers have narrow slits or furrows that run lengthwise along the anthers
- Anther spurs
- the anthers do not have spurs on them
- Calyx symmetry
- there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
- Carpel hairs
- the carpels have no hairs
- Carpels fused
- the carpels are fused to one another
- Cilia on petals
-
- the petal margins do not have cilia
- the petal margins have cilia
- Cleistogamous flowers
- there are no cleistogamous flowers on the plan
- Corolla palate
- no
- Corona lobe length
- 0 mm
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Filament length
- 0.5–1 mm
- Filament surface
- the filament is smooth, with no hairs or scales
- Flower description
- the flower has a superior ovary, and lacks a hypanthium
- Flower diameter
- 0.8–2 mm
- Flower length
- 3–6.2 mm
- Flower orientation
- the flower points upwards or is angled outwards
- Flower petal color
-
- pink to red
- white
- Flower reproductive parts
- the flower has both pollen- and seed-producing parts
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Flowers sunken into stem
- no
- Form of style
- the style is lobed at the tip, and unbranched
- Fused stamen clusters
- NA
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Hairs on flower stalk
- the flower stalk has no hairs on it
- Hairs on inflorescence
- the axis of the inflorescence has no hairs on it
- Horns in hoods (Asclepias)
- NA
- Hypanthium
- the flower does not have a hypanthium
- Inflorescence one-sided
- the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
- Inner tepals (Rumex)
- NA
- Interior flower disk
- NA
- Marks on petals
- there are no noticeable marks on the petals
- Nectar spur
- the flower has no nectar spurs
- Number of carpels
- 2
- Number of sepals, petals or tepals
- there are four petals, sepals, or tepals in the flower
- Number of styles
- 1
- Ovary position
- the ovary is above the point of petal and/or sepal attachment
- Perianth shape
- the perianth is campanulate (bell-shaped, with a tube about as long as wide, flaring at the mouth)
- Petal and sepal arrangement
- the flower includes two cycles of petal- or sepal-like structures
- Petal and sepal colors
-
- pink to red
- white
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal folding in bud
- the petals in bud are arranged in a cycle with edges overlapping like roof shingles (imbricate)
- Petal folds or pleats
- the petals of the flower do not have folds or plaits
- Petal glandular dots or scales
- no
- Petal hairs (Viola)
- NA
- Petal length
- 3–6.2 mm
- Petal length relative to sepals
- the petals are about equal in length to the sepals
- Petal number
- 4
- Petal shape
-
- the petal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)
- the petal outline is oblong (rectangular, but with rounded ends)
- Petal tip shape
- the petal tip is acute (sharply pointed)
- Petal tips (Cuscuta)
- NA
- Petal width
- 0.8–2 mm
- Raceme attachment (Veronica)
- NA
- Reproductive system
- all the flowers have both carpels and stamens (synoecious)
- Scales inside corolla
- no
- Sepal and petal color
- the sepals are different from the petals
- Sepal appearance
- the sepals are green or brown, and leaf-like in texture
- Sepal appendages
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Sepal auricles
- the sepals have no auricles
- Sepal cilia
-
- the sepals do not have cilia
- the sepals have cilia on their edges
- Sepal color
-
- blue to purple
- green to brown
- Sepal features
- one or more winged
- Sepal length
- 1.5–3.2 mm
- Sepal number
- 4
- Sepal orientation
- the sepals are pressed against the corolla, or jutting stiffly upward
- Sepal relative length
- the sepal lobes are longer than the fused portion
- Sepal shape
-
- the sepal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)
- the sepal outline is subulate (narrowly tapering from the base to the apex; awl-shaped)
- Sepal texture
- the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
- Sepal tip shape
-
- the sepal tip is acuminate (tapers to a very narrow point)
- the sepal tip is acute (is sharply pointed)
- Sepal uniformity
- all the sepals are about the same size
- Sepals fused only to sepals
- the sepals are fused to each other (not other flower parts), at least near their bases
- Spur length
- 0 mm
- Spur number
- NA
- Stamen appendages
- stamen appendages are absent
- Stamen attachment
- the stamens are attached at or near the bases of the petals or tepals
- Stamen morphology
- the stamens within each cycle are the same
- Stamen number
- 4
- Stamen position relative to petals
- the stamens are lined up with the sepals
- Stamen relative length
- anything
- Stamens fused
- the stamens are not attached to one another
- Staminodes
- there are no staminodes on the flower
- Stigma length
- 0.8–1.5 mm
- Stigma position
- the stigmas are positioned in lines on the ovary
- Style petal-like
- the styles are not petal-like
- Style relative length
- the stigma does not protrude beyond the mouth of the corolla
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene relative orientation
- NA
- Achene shape
- NA
- Achene surface (Polygonum)
- NA
- Achene type
- NA
- Berry color
- NA
- Capsule color (Viola)
- NA
- Capsule splitting
- the capsule splits by two main valves, teeth or pores
- Fruit (pyxis) dehiscence
- NA
- Fruit cross-section
- the fruit is round in cross-section
- Fruit features (Brassicaceae)
- NA
- Fruit length
- 4–6 mm
- Fruit length relative to sepals
- the fruit is longer than its associated sepals
- Fruit locules
- one
- Fruit shape
- the fruit is ovoid (egg-shaped)
- Fruit stalk orientation
- the fruits point upward or spread or curve outward
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit type (specific)
- the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)
- Legumes (Fabaceae)
- NA
- Mericarp length
- 0 mm
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
- NA
- Placenta arrangement
- the plant has parietal placentation, where ovules develop on the wall or slight outgrowths of the wall forming broken partitions within a compound ovary
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- NA
- Schizocarpic fruit segments
- 0
- Seed length
- 0.15–0.1 mm
- Seed relative length
- the seed is about as long as it is wide
- Septum in fruit (Brassicaceae)
- NA
- Wings on fruit
- the fruit does not have wings on it
- prickles on fruits
- the fruits do not have thorn-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades do not have glandular dots or scales
- Sap
- the sap is clear and watery
- Sap color
- the sap is clear
-
Growth form
- Growth form
- the plant is an herb (it has self-supporting stems)
- Lifespan
-
- the plant lives for two years
- the plant lives only a single year or less
- Parasitism
- the plant is a parasite of fungi (mycoheterotrophic)
- Plant color
- the plant lacks green pigments, even on the leaves or young stems
- Plants darken when dry
- no
- Spines on plant
- the plant has no spines
-
Leaves
- Bracteole edges
- NA
- Bracteole length
- 0 mm
- Bracteole number (Apiaceae)
- 0
- Bracteole shape
- NA
- Bracteoles
- there are no bracteoles on the plant
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length (compound lvs only)
- 0 mm
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Final leaf segment width (compound lvs only)
- 0 mm
- Floral bract texture
- NA
- Floral bracts
- the flower does not have bracts
- Flower bract length
- 0 mm
- Hairs on leaf stalk
- NA
- Hairs on underside of leaf
- the underside of the leaf is not hairy, or it has very few hairs
- Hairs on upper side of leaf
- the upper side of the leaf is not hairy, or it has very few hairs
- Hooked hairs on underside of leaf
- no
- Leaf arrangement
- alternate: there is one leaf per node along the stem
- Leaf blade bloom
- NA
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf blade flatness
- NA
- Leaf blade hairs
- NA
- Leaf blade length
- 0.5–4.5 mm
- Leaf blade primary vein pattern
- the secondary veins branch off at intervals from the primary vein
- Leaf blade shape
- the leaf blade is subulate (awl-shaped; narrowly tapering from the base to the tip)
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf blade vein pattern
- NA
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are small and thin and lack leaf stalks
- Leaf hair orientation
- NA
- Leaf spines
- there are no spines on the leaf edges
- Leaf stalk
- the leaves have no leaf stalks, but attach directly to the stem
- Leaf stalk attachment to leaf
- NA
- Leaf stalk base
- NA
- Leaf teeth and lobes
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf tip
- the tip of the leaf blade is acuminate (tapers to a long, thin point)
- Leaf tufts in axils
- there are no clusters of smaller leaves growing out of axils
- Leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Leaf types
- There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).
- Leaf variation
- the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
- there is one leaf per node along the stem
- Pinnately compound leaf type
- NA
- Specific leaf type
- the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
- Stipels
- NA
- Stipule edges
- NA
- Stipule features
- NA
- Stipule fused to leaf stalk
- NA
- Stipule length
- 0 mm
- Stipule shape
- NA
- Stipules
- there are no stipules on the plant
- Teeth per side of leaf blade
- 0
-
Place
- Habitat
-
- terrestrial
- wetlands
- Specific habitat
-
- bogs
- fens
- man-made or disturbed habitats
- meadows or fields
- swamps
-
Scent
- Plant odor
- the plant does not have much of a smell
-
Stem, shoot, branch
- Branched tendrils
- NA
- Direction of stem hairs
- NA
- Flowering stem cross-section
-
- the flowering stem has strong or sharp angles, but is not square
- the flowering stem is circular, or with lots of small angles so that it is roughly circular
- Hair between stem nodes
- the stem has no hairs between the nodes
- Hairs between stem nodes
- NA
- Hooked hairs on stem between nodes
- no
- Leaves on stem
- there is at least one full leaf above the base of the flowering stem
- Plant height
- 3–25 cm
- Stem hair distribution
- NA
- Stem orientation
- the stems are upright or angled outwards
- Stem roughness between nodes
- the stem does not feel rough
- Stem spacing
- the plant is solitary, or a few plants are growing together
- Stem succulence
- the stems are not succulent
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
- Wings on stem
- the stem does not have wings on it
Wetland status
Occurs only in wetlands. (Wetland indicator code: OBL)
In New England
Distribution
- Connecticut
- present
- Maine
- present
- Massachusetts
- present
- New Hampshire
- present
- Rhode Island
- present
- Vermont
- absent
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- New Hampshire
- historical (S-rank: SH), endangered (code: E)
- Rhode Island
- extremely rare (S-rank: S1), concern (uncertain) (code: C*)
From Flora Novae Angliae dichotomous key
1. Bartonia iodandra B. L. Robins. NC
purple screwstem. Bartonia paniculata (Michx.) Muhl. var. intermedia Fern.; B. paniculata (Michx.) Muhl. ssp. iodandra (B.L. Robins.) J. Gillett; B. paniculata (Michx.) Muhl. var. iodandra (B.L. Robins.) Fern.; B. paniculata (Michx.) Muhl. var. sabulonensis (Fern.) Fern.; B. virginica (L.) B.S.P. var. sabulonensis (Fern.) Boivin • MA, ME, NH, RI. Swamps, peatlands, abandoned borrow pits, meadows. Bartonia iodandra has been treated as a subspecies or variety of B. paniculata by most authors, likely due to similarities in leaf arrangement and stigma length. However, it possesses many character states that are transitional to B. virginica, including stem stature and coloration, petal shape and apex, anther apex, stamen color, and seed size. Gillett (1959) noted its probable hybrid origin. As such, it is inappropriate to ally it with one of its putative parents. Its occurrence in Newfoundland (beyond the range of its parents) and production of well-formed seeds support its recognition as a hybrid-derived species (rather than as an F₁ hybrid).
Native to North America?
Yes
Sometimes confused with
- Bartonia paniculata:
- petals lanceolate and acute at the apex, anthers usually yellow and mostly 0.3-0.5 mm long, and stems often slender, lax, and green (vs. B. iodandra, with petals oblong-lanceolate and acute at the apex or infrequently with an obscure apiculus, anthers usually purple and mostly 0.5-0.6 mm long, and stems stouter, strict, usually purple or purple tinged).
- Bartonia virginica:
- leaves opposite and stigmas mostly 1.5-2.3 mm long (vs. B. iodandra, with leaves alternate and stigmas 0.8-1.5 mm long).
Synonyms
- Bartonia paniculata (Michx.) Muhl. ssp. iodandra (B.L. Robins.) J. Gillett
- Bartonia paniculata (Michx.) Muhl. var. intermedia Fern.
- Bartonia paniculata (Michx.) Muhl. var. iodandra (B.L. Robins.) Fern.
- Bartonia paniculata (Michx.) Muhl. var. sabulonensis (Fern.) Fern.
- Bartonia virginica (L.) B.S.P. var. sabulonensis (Fern.) Boivin