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Plantago rugelii — Rugel's plantain

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Facts

Rugel's plantain is a native plantain of disturbed areas throughout New England. You can recognize it by its leaves, which are purplish along the petiole near where they join the stem. The species is named for the German-born botanist and physician, Ferdinand Rugel (1806-1879), who collected many new species in the Appalachian and southern regions of the young United States, as well as Cuba.

Habitat

Anthropogenic (man-made or disturbed habitats), grassland, meadows and fields

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
terrestrial
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Flower petal color
  • other
  • white
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
basal: the leaves are growing only at the base of the plant
Leaf blade edges
the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
Number of sepals, petals or tepals
  • there are four petals, sepals, or tepals in the flower
  • there are three petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
4
Fruit type (general)
the fruit is dry and splits open when ripe
Fruit length
3–6 mm
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Anther spurs
    the anthers do not have spurs on them
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Filament surface
    the filament is smooth, with no hairs or scales
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower petal color
    • other
    • white
    Flower reproductive parts
    • the flower has both pollen- and seed-producing parts
    • the flower has either only pollen- or only seed-producing parts
    Flower symmetry
    there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is knob-like at the tip, and unbranched
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inner tepals (Rumex)
    NA
    Nectar spur
    the flower has no nectar spurs
    Number of pistils
    • 1
    • 2
    Number of sepals, petals or tepals
    • there are four petals, sepals, or tepals in the flower
    • there are three petals, sepals, or tepals in the flower
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    • other
    • white
    Petal appearance
    the petals are green and/or leafy in texture
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal number
    4
    Petal tips (Cuscuta)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal number
    4
    Stamen attachment
    the stamens are attached at or near the bases of the petals or tepals
    Stamen number
    4
    Stamen position relative to petals
    the stamens are lined up with the sepals
    Staminodes
    there are no staminodes on the flower
    Stigma position
    the stigmas are positioned at the tip of the style
    Style petal-like
    the styles are not petal-like
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    NA

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    NA
    Capsule splitting
    NA
    Fruit (pyxis) dehiscence
    the pyxis splits below the middle
    Fruit features (Brassicaceae)
    NA
    Fruit length
    3–6 mm
    Fruit locules
    two
    Fruit type (general)
    the fruit is dry and splits open when ripe
    Fruit type (specific)
    the fruit is a pyxis (when dry it splits around the middle, and the top falls off, exposing the seeds)
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines

  • Leaves

    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    • the bracts are smooth and hairless
    • the bracts have an obvious ridge at the center (at least near the base)
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Hairs on underside of leaf
    the underside of the leaf is fuzzy or hairy
    Leaf arrangement
    basal: the leaves are growing only at the base of the plant
    Leaf blade base
    the leaf has a distinct leaf stalk (petiole)
    Leaf blade base shape
    • the base of the leaf blade is cordate (heart-shaped, has rounded lobes at the base)
    • the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
    Leaf blade edges
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade length
    40–180 mm
    Leaf blade shape
    • the leaf blade is cordate (heart-shaped with backward-facing rounded lobes), or sagittate (arrow-shaped with backward-facing pointed lobes)
    • the leaf blade is elliptic (widest near the middle and tapering at both ends)
    • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have leaf stalks
    Leaf teeth and lobes
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf variation
    NA
    Leaflet number
    0
    Leaves per node
    the leaves are growing only at the base of the plant (i.e., basal)
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule features
    NA
    Stipules
    there are no stipules on the plant

  • Place

    Habitat
    terrestrial
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • grasslands
    • man-made or disturbed habitats
    • meadows or fields

  • Scent

    Plant odor
    the plant does not have much of a smell

  • Stem, shoot, branch

    Branched tendrils
    NA
    Hair between stem nodes
    the stem has hairs between the nodes
    Hairs between stem nodes
    the hairs on the stem are plain, without glands or branches, and not tangled
    Hooked hairs on stem between nodes
    NA
    Leaves on stem
    the flowering stem has no leaves above the base
    Plant height
    5–30 cm
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils

Wetland status

Occurs in wetlands or non-wetlands. (Wetland indicator code: FAC)

In New England

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
present
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

Massachusetts
widespread (S-rank: S5)

From Flora Novae Angliae dichotomous key

11.  Plantago rugelii Dcne. N

Rugel’s plantain. Plantago rugelii Dcne. var. asperula Farw. • CT, MA, ME, NH, RI, VT. Fields, roadsides, waste areas, lawns, trail edges.

Native to North America?

Yes

Sometimes confused with

Plantago intermedia:
pyxis mostly 2.5–4 mm tall, circumscissile near the middle, containing mostly 14–22 seeds, bracts broad-ovate, and seeds 1–1.7 mm long, reticulate-patterned (vs. P. rugelii, with the pyxis mostly 4–6 mm tall, circumscissile near the base, containing mostly 4–9 seeds, bracts narrow-triangular, and seeds mostly 1.5–2 mm long, not reticulate-patterned).
Plantago major:
pyxis mostly 2.5–4 mm tall, circumscissile near the middle, containing mostly 6–11 seeds, bracts broad-ovate, and seeds 1–1.7 mm long, reticulate-patterned (vs. P. rugelii, with the pyxis mostly 4–6 mm tall, circumscissile near the base, containing mostly 4–9 seeds, bracts narrow-triangular, and seeds mostly 1.5–2 mm long, not reticulate-patterned).

Synonyms

  • Plantago rugelii Dcne. var. asperula Farw.

Family

Plantaginaceae

Genus

Plantago