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Polygonum achoreum — leathery knotweed

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Facts

Leathery knotweed is found in disturbed sites, field edges and even sidewalks. Plants are often covered with a whitish powdery mildew.

Habitat

Anthropogenic (man-made or disturbed habitats), meadows and fields

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
terrestrial
New England state
  • Maine
  • Vermont
Flower petal color
  • green to brown
  • pink to red
  • yellow
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
alternate: there is one leaf per node along the stem
Leaf blade edges
the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
Number of sepals, petals or tepals
there are five petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
  • 5
  • 6
  • 7
  • 8
Fruit type (general)
the fruit is dry but does not split open when ripe
Fruit length
2.4–3.5 mm
Show all characteristics
  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located
  • Flowers

    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Anther spurs
    the anthers do not have spurs on them
    Calyx growth after flowering
    the calyx does not grow to cover or partially cover the fruit
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Filament surface
    the filament is smooth, with no hairs or scales
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower length
    2.5–3.7 mm
    Flower number
    1–5
    Flower petal color
    • green to brown
    • pink to red
    • yellow
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
    Flowers sunken into stem
    no
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    • the flower does not have a hypanthium
    • the flower has a hypanthium
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inner tepals (Rumex)
    NA
    Length of flower stalk
    1.3–2 mm
    Length of peduncle
    0 mm
    Marks on petals
    there are no noticeable marks on the petals
    Nectar spur
    the flower has no nectar spurs
    Number of branches in umbel
    0
    Number of carpels
    3
    Number of pistils
    1
    Number of sepals, petals or tepals
    there are five petals, sepals, or tepals in the flower
    Number of styles
    3
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Perianth shape
    • the perianth is campanulate (bell-shaped, with a tube about as long as wide, flaring at the mouth)
    • the perianth is urceolate (urn-shaped, with a swollen tube contracted near the top, then slightly expanded in a narrow rim)
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    • green to brown
    • pink to red
    • yellow
    Petal appearance
    • the petals are green and/or leafy in texture
    • the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folds or pleats
    • the petals of the flower do not have folds or plaits
    • the petals of the flower have folds or plaits on them
    Petal hairs (Viola)
    NA
    Petal hairs on inner/upper surface
    there are no hairs on the inner/upper petal surface
    Petal length
    2.6–4 mm
    Petal nectaries
    the petals do not have nectaries
    Petal number
    2
    Petal shape
    the petal outline is oblong (rectangular, but with rounded ends)
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the petals and sepals are similar
    Sepal appearance
    the sepals are green or brown, and leaf-like in texture
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal color
    • green to brown
    • yellow
    Sepal features
    • one or more sepals are arched and enfolding, hood-shaped
    • one or more sepals have a prominent keel (a ridge that runs along the midline from base to tip)
    Sepal length
    2.6–4 mm
    Sepal number
    3
    Sepal orientation
    the sepals are pressed against the corolla, or jutting stiffly upward
    Sepal relative length
    • the sepal lobes are approximately the same length as the fused portion
    • the sepal lobes are longer than the fused portion
    • the sepal lobes are shorter than the fused portion
    Sepal shape
    the sepal outline is oblong (rectangular, but with rounded ends)
    Sepal texture
    the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
    Sepal tip shape
    the sepal tip is rounded
    Sepal uniformity
    all the sepals are about the same size
    Sepals fused only to sepals
    the sepals are fused to each other (not other flower parts), at least near their bases
    Spur length
    0 mm
    Spur number
    NA
    Stamen morphology
    the stamens within a cycle differ in length or width
    Stamen number
    • 5
    • 6
    • 7
    • 8
    Stamens fused
    the stamens are not attached to one another
    Staminodes
    there are no staminodes on the flower
    Style petal-like
    the styles are not petal-like
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    NA
  • Fruits or seeds

    Achene relative orientation
    the achenes are perpendicular to the plane of the perianth (vertical)
    Achene shape
    the achenes are another shape in outline
    Achene surface (Polygonum)
    the achene has tiny rounded bumps or projections on it (papillose)
    Achene type
    the fruit is an achene (dry, indehiscent, and usually one-seeded)
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    NA
    Capsule splitting
    NA
    Carpel beak length
    0 mm
    Fruit (pyxis) dehiscence
    NA
    Fruit cross-section
    the fruit is roughly triangular in cross-section
    Fruit features (Brassicaceae)
    NA
    Fruit length
    2.4–3.5 mm
    Fruit length relative to sepals
    the fruit is shorter than its associated sepals
    Fruit locules
    one
    Fruit shape
    • the fruit is lanceoloid (narrow, widest below the middle and tapering at both ends)
    • the fruit is ovoid (egg-shaped)
    Fruit stalk orientation
    the fruits point upward or spread or curve outward
    Fruit type (general)
    the fruit is dry but does not split open when ripe
    Fruit type (specific)
    the fruit is an achene (dry, usually one-seeded, does not separate or split open at maturity)
    Hair type on fruit
    NA
    Hairs on fruit
    the fruits are not hairy
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Ovary stipe length
    0 mm
    Placenta arrangement
    the plant has basal placentation, where one or a few ovules develop at the base of a simple or compound ovary
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Seed length
    1 mm
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures
  • Glands or sap

    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear
  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives only a single year or less
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    there is a thickened taproot on the plant
  • Leaves

    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bract texture
    NA
    Floral bracts
    the flower does not have bracts
    Flower bract length
    0 mm
    Hairs on underside of leaf
    the underside of the leaf is not hairy, or it has very few hairs
    Hairs on upper side of leaf
    the upper side of the leaf is not hairy, or it has very few hairs
    Hooked hairs on underside of leaf
    no
    Inflated hairs on leaf
    the leaf blade does not have inflated hairs on it
    Leaf arrangement
    alternate: there is one leaf per node along the stem
    Leaf blade base
    the leaf has a distinct leaf stalk (petiole)
    Leaf blade base symmetry
    the leaf blade base is symmetrical
    Leaf blade bloom
    • the underside of the leaf has no noticeable bloom
    • there is a noticeable powdery or waxy bloom on the underside of the leaf
    Leaf blade edges
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade flatness
    the leaf is flat (planar) at the edges
    Leaf blade hairs
    NA
    Leaf blade length
    8–35 mm
    Leaf blade primary vein pattern
    the secondary veins branch off at intervals from the primary vein
    Leaf blade shape
    • the leaf blade is elliptic (widest near the middle and tapering at both ends)
    • the leaf blade is obovate (egg-shaped, but with the widest point above the middle of the leaf blade)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade vein pattern
    the major veins of the leaf blade branch, but do not rejoin
    Leaf blade veins
    the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)
    Leaf blade width
    3–15 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf hair orientation
    NA
    Leaf shiny
    the upper side of the leaf is dull or slightly shiny
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have leaf stalks
    Leaf stalk attachment to leaf
    the petiole attaches at the basal margin of the leaf blade
    Leaf stalk length
    0.3–1.5 mm
    Leaf teeth and lobes
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf tip
    the tip of the leaf blade is rounded, with no point
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf types
    NA
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaflet petiolules
    NA
    Leaves per node
    there is one leaf per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule edges
    • the stipule margins do not have teeth
    • the stipule margins have teeth on them
    Stipule features
    the stipules have two lobes at their tips
    Stipule fused to leaf stalk
    the stipules are fused to the petioles for some or most of their length
    Stipule length
    5–12 mm
    Stipule shape
    • the stipules are capillary (very fine and hair-like, not flattened)
    • the stipules are tubular (cylindrical and hollow)
    Stipules
    the plant has stipules
    Teeth per side of leaf blade
    0
  • Place

    Habitat
    terrestrial
    New England state
    • Maine
    • Vermont
    Specific habitat
    • man-made or disturbed habitats
    • meadows or fields
  • Scent

    Plant odor
    the plant does not have much of a smell
  • Stem, shoot, branch

    Branched tendrils
    NA
    Direction of stem hairs
    NA
    Flowering stem cross-section
    the flowering stem is circular, or with lots of small angles so that it is roughly circular
    Hair between stem nodes
    the stem has no hairs between the nodes
    Hairs between stem nodes
    the stem has no hairs between the nodes
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Length of hairs between stem nodes
    0 mm
    Plant height
    50–70 cm
    Stem bloom
    there is no powdery or waxy film on the stem
    Stem hair distribution
    NA
    Stem nodes swollen
    the stem is swollen at the nodes
    Stem orientation
    • the stems are upright or angled outwards
    • the stems trail at the base, but may turn upwards at the tips
    Stem roughness between nodes
    NA
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils
    Wings on stem
    the stem does not have wings on it

Wetland status

Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)

In New England

Distribution

Connecticut
absent
Maine
present
Massachusetts
absent
New Hampshire
absent
Rhode Island
absent
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

New Hampshire
unrankable (S-rank: SU), Ind (code: Ind)
Vermont
uncommon (S-rank: S3)

From Flora Novae Angliae dichotomous key

1.  Polygonum achoreum Blake N

leathery knotweed. Polygonum erectum L. ssp. achoreum (Blake) A. & D. Löve • ME, VT. Fields, roadsides, waste areas, railroads, lawns. The report of this species in CT by Magee and Ahles (1999) is based on a specimen of Polygonum buxiformeAhles 85330 ( MASS!).

Native to North America?

Yes

Sometimes confused with

Polygonum ramosissimum:
petal-like parts of the flower fused at the base for 20-38% of their length and leaf blades lanceolate to narrow-elliptic (vs. P. achoreum, with petal-like parts of the flower fused at the base for 40-55% of their length and leaf blades mostly elliptic to obovate).

Synonyms

  • Polygonum erectum ssp. achoreum (Blake) A. & D. Löve

Family

Polygonaceae

Genus

Polygonum