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Physalis heterophylla — clammy ground-cherry

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Facts

A native perennial, clammy ground-cherry fruits are edible when ripe, but the rest of the plant is toxic.

Habitat

Anthropogenic (man-made or disturbed habitats), meadows and fields

New England distribution

Adapted from BONAP data

Found this plant? Take a photo and post a sighting.

North America distribution

Adapted from BONAP data

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Characteristics

Habitat
terrestrial
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Flower petal color
yellow
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
alternate: there is one leaf per node along the stem
Leaf blade edges
  • the edge of the leaf blade has lobes, or it has both teeth and lobes
  • the edge of the leaf blade has teeth
  • the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
Number of sepals, petals or tepals
there are five petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
5
Fruit type (general)
the fruit is fleshy
Fruit length
10 mm
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Anther length
    3.5–4.5 mm
    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Anther spurs
    the anthers do not have spurs on them
    Calyx growth after flowering
    the calyx grows to cover or partially cover the fruit
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower length
    10–20 mm
    Flower number
    1
    Flower orientation
    the flower bends downwards or hangs downwards
    Flower petal color
    yellow
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
    Flowers sunken into stem
    no
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Hairs on flower stalk
    the flower stalk has hairs on it
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Hypanthium length
    0 mm
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inner tepals (Rumex)
    NA
    Length of flower stalk
    Up to 30 mm
    Length of peduncle
    0 mm
    Marks on petals
    the petals have spots or streaks on them
    Nectar spur
    the flower has no nectar spurs
    Number of branches in umbel
    0
    Number of carpels
    2
    Number of pistils
    1
    Number of sepals, petals or tepals
    there are five petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Perianth shape
    • the perianth is campanulate (bell-shaped, with a tube about as long as wide, flaring at the mouth)
    • the perianth is rotate (platter-shaped, the corolla flattened, circular, with nearly horizontally spreading lobes)
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    yellow
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folding in bud
    • the petals are arranged some other way in bud
    • the petals in bud meet exactly at the margins without overlapping (valvate)
    Petal hairs (Viola)
    NA
    Petal length
    10–20 mm
    Petal length relative to sepals
    the petals are longer than the sepals
    Petal number
    5
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal auricles
    the sepals have no auricles
    Sepal color
    • green to brown
    • yellow
    Sepal length
    4–15 mm
    Sepal number
    5
    Sepal relative length
    the sepal lobes are longer than the fused portion
    Sepal shape
    • the sepal outline is ovate (widest below the middle and broadly tapering at both ends)
    • the sepal outline is roughly triangular
    Sepal texture
    • the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
    • the sepals are thin and dry, paperlike
    Sepals fused only to sepals
    the sepals are fused to each other (not other flower parts), at least near their bases
    Stamen attachment
    the stamens are attached at or near the bases of the petals or tepals
    Stamen lengths differ
    the stamens are all approximately the same length
    Stamen morphology
    the stamens within each cycle are the same
    Stamen number
    5
    Stamen position relative to petals
    the stamens are lined up with the sepals
    Stamen relative length
    anything
    Stamens fused
    the stamens are not attached to one another
    Staminodes
    there are no staminodes on the flower
    Style petal-like
    the styles are not petal-like
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    NA

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    green
    Capsule color (Viola)
    NA
    Capsule ribs
    NA
    Capsule splitting
    NA
    Carpel beak length
    0 mm
    Fruit (pyxis) dehiscence
    NA
    Fruit beak length
    0 mm
    Fruit cross-section
    the fruit is round in cross-section
    Fruit features (Brassicaceae)
    NA
    Fruit length
    10 mm
    Fruit length relative to sepals
    the fruit is shorter than its associated sepals
    Fruit locules
    two
    Fruit shape
    the fruit is spherical
    Fruit stalk orientation
    the fruits curve or droop downwards
    Fruit type (general)
    the fruit is fleshy
    Fruit type (specific)
    the fruit is a berry (fleshy, with the wall enclosing one or more sections, with two or more seeds)
    Fruit width
    10 mm
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    the ripe berries are mostly one color without spots or streaks
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Seed length
    2–2.5 mm
    Seed surface
    the seed has pits or depressions on it
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    the plant has a rhizome (a horizontal underground stem with roots growing from it)

  • Leaves

    Bracteole edges
    NA
    Bracteole length
    0 mm
    Bracteole number (Apiaceae)
    0
    Bracteole shape
    NA
    Bracteoles
    there are no bracteoles on the plant
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Hairs on leaf stalk
    the petiole has hairs on it
    Hairs on underside of leaf
    the underside of the leaf is fuzzy or hairy
    Hairs on upper side of leaf
    the upper side of the leaf is fuzzy or hairy
    Hooked hairs on underside of leaf
    no
    Inflated hairs on leaf
    the leaf blade does not have inflated hairs on it
    Leaf arrangement
    alternate: there is one leaf per node along the stem
    Leaf blade base
    the leaf has a distinct leaf stalk (petiole)
    Leaf blade base shape
    • the base of the leaf blade is cordate (heart-shaped, has rounded lobes at the base)
    • the base of the leaf blade is rounded
    Leaf blade base symmetry
    the leaf blade base is symmetrical
    Leaf blade bloom
    the underside of the leaf has no noticeable bloom
    Leaf blade edges
    • the edge of the leaf blade has lobes, or it has both teeth and lobes
    • the edge of the leaf blade has teeth
    • the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade hairs
    the leaf blade has simple hairs with no glands, and not tangled or wooly
    Leaf blade length
    30–80 mm
    Leaf blade primary vein pattern
    the secondary veins branch off at intervals from the primary vein
    Leaf blade shape
    • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
    • the leaf blade is rhombic (roughly diamond-shaped)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade texture
    the leaf blade is scarious (thin and dry like paper) or membranaceous (thin and flexible and almost translucent like a membrane)
    Leaf blade vein pattern
    the major veins of the leaf blade branch, but do not rejoin
    Leaf blade veins
    the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)
    Leaf blade width
    30–80 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf hair orientation
    the hairs are standing up straight or curved in different directions
    Leaf shiny
    the upper side of the leaf is dull or slightly shiny
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have leaf stalks
    Leaf stalk attachment to leaf
    the petiole attaches at the basal margin of the leaf blade
    Leaf teeth and lobes
    • the edge of the leaf blade is entire (has no teeth or lobes)
    • the leaf blade margin has outward-pointing teeth
    Leaf tip
    the tip of the leaf blade is acute (sharply pointed)
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf types
    • NA
    • There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaflet petiolules
    NA
    Leaves per node
    there is one leaf per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule edges
    NA
    Stipule features
    NA
    Stipule fused to leaf stalk
    NA
    Stipule length
    0 mm
    Stipule shape
    NA
    Stipules
    there are no stipules on the plant

  • Place

    Habitat
    terrestrial
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • man-made or disturbed habitats
    • meadows or fields

  • Stem, shoot, branch

    Branched tendrils
    NA
    Direction of stem hairs
    the hairs point mostly upwards to outwards
    Flowering stem cross-section
    the flowering stem is circular, or with lots of small angles so that it is roughly circular
    Hair between stem nodes
    the stem has hairs between the nodes
    Hairs between stem nodes
    at least some of the hairs on the stem have glands
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    20–90 cm
    Stem bloom
    there is no powdery or waxy film on the stem
    Stem hair distribution
    the hairs on the stem are distributed more of less uniformly
    Stem nodes swollen
    the stem is not swollen at the nodes
    Stem orientation
    the stems are upright or angled outwards
    Stem roughness between nodes
    the stem does not feel rough
    Stem succulence
    the stems are not succulent
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils
    Wings on stem
    the stem does not have wings on it

Wetland status

Not classified

In New England

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
present
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

var. heterophylla

Massachusetts
fairly widespread (S-rank: S4)

Subspecies and varieties

Our variety is Physalis heterophylla Nees var. heterophylla.

From Flora Novae Angliae dichotomous key

4.  Physalis heterophylla Nees var. heterophylla N

clammy ground-cherry. Physalis ambigua (Gray) Britt.; P. heterophylla Nees var. ambigua (Gray) Rydb.; P. heterophylla Nees var. clavipes Fern.; P. heterophylla Nees var. nyctaginea (Dunal) Rydb.; P. heterophylla Nees var. villosa Waterfall; P. nyctaginea Dunal; P. sinuata Rydb. • CT, MA, ME, NH, RI, VT. Fields, roadsides, waste areas.

Native to North America?

Yes and no (some introduced)

Sometimes confused with

Physalis grisea:
petals 4-10 mm long, sepals mostly 3-7 mm long during flowering, and plants annual from a taproot (vs. P. heterophylla, with petals 10-20 mm long, sepals mostly 6-12 mm long during flowering, and plants perennial from rhizomes).
Physalis longifolia:
apical half of the stem pubescent entirely or nearly entirely with eglandular hairs of various lengths, these often short, stiff, usually appressed hairs, and leaf blades narrow-lanceolate to ovate, narrow-cuneate to broad-cuneate at the base (vs. P. heterophylla, with the apical half of the stem pubescent and often other parts of the plant pubescent with stipitate-glands intermixed with short, eglandular hairs and longer, eglandular, multicellular hairs, and leaf blades ovate to rhombic, broad-rounded to cordate at the base).

Synonyms

  • Physalis ambigua (Gray) Britt.
  • Physalis heterophylla var. ambigua (Gray) Rydb.
  • Physalis heterophylla var. clavipes Fern.
  • Physalis heterophylla var. nyctaginea (Dunal) Rydb.
  • Physalis heterophylla var. villosa Waterfall
  • Physalis nyctaginea Dunal
  • Pysalis sinuata Rydb.

Family

Solanaceae

Genus

Physalis