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Lycopus americanus — American water-horehound

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Facts

The Meskwaki used American water-horehound as an analgesic and gastro-intestinal aid.

Habitat

Shores of rivers or lakes, swamps, wetland margins (edges of wetlands)

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
  • terrestrial
  • wetlands
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Flower petal color
white
Leaf type
the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
Leaf arrangement
opposite: there are two leaves per node along the stem
Leaf blade edges
  • the edge of the leaf blade has lobes, or it has both teeth and lobes
  • the edge of the leaf blade has teeth
Flower symmetry
  • there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
  • there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
Number of sepals, petals or tepals
there are four petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
1 or 2
Fruit type (general)
the fruit is dry but does not split open when ripe
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Anther length
    0.26–0.5 mm
    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Cleistogamous flowers
    there are no cleistogamous flowers on the plan
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Filament surface
    the filament is smooth, with no hairs or scales
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower petal color
    white
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    • there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
    • there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is lobed at the tip, and unbranched
    Fringed petal edges
    the petals are not fringed
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inner tepals (Rumex)
    NA
    Interior flower disk
    the flower has an interior disc
    Marks on petals
    • the petals have spots or streaks on them
    • there are no noticeable marks on the petals
    Nectar spur
    the flower has no nectar spurs
    Number of branches in umbel
    0
    Number of carpels
    2
    Number of sepals, petals or tepals
    there are four petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Perianth shape
    the perianth is campanulate (bell-shaped, with a tube about as long as wide, flaring at the mouth)
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    white
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal hairs on inner/upper surface
    there are hairs on the inner/upper petal surface
    Petal length
    1.8–4.4 mm
    Petal length relative to sepals
    the petals are shorter than the sepals
    Petal number
    4
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appearance
    the sepals are green or brown, and leaf-like in texture
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal color
    green to brown
    Sepal length
    2–3.3 mm
    Sepal number
    5
    Sepal relative length
    the sepal lobes are longer than the fused portion
    Sepal shape
    the sepal outline is roughly triangular
    Sepal texture
    the sepals are firm or rigid
    Sepal tip shape
    the sepal tip is acuminate (tapers to a very narrow point)
    Sepals fused only to sepals
    the sepals are fused to each other (not other flower parts), at least near their bases
    Spur length
    0 mm
    Spur number
    NA
    Stamen attachment
    the stamens are not attached to the petals or tepals
    Stamen number
    1 or 2
    Stamen relative length
    anything
    Stamens fused
    the stamens are not attached to one another
    Staminodes
    • the flower has staminodes
    • there are no staminodes on the flower
    Style relative length
    the stigma protrudes beyond the mouth of the corolla
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    • NA
    • the upper lip of the bilabiate corolla has one lobe

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    NA
    Capsule splitting
    NA
    Fruit (pyxis) dehiscence
    NA
    Fruit beak length
    0 mm
    Fruit features (Brassicaceae)
    NA
    Fruit length relative to sepals
    the fruit is shorter than its associated sepals
    Fruit locules
    four
    Fruit type (general)
    the fruit is dry but does not split open when ripe
    Fruit type (specific)
    the fruit is a schizocarp (when dry it splits into sections, each holding one or more seeds)
    Legumes (Fabaceae)
    NA
    Mericarp length
    1–1.4 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    the fruit is not flattened
    Schizocarpic fruit segments
    4
    Seed number
    2
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives more than two years
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    the plant has a rhizome (a horizontal underground stem with roots growing from it)

  • Leaves

    Bracteole number (Apiaceae)
    0
    Bracteoles
    the plant has bracteoles between the primary bracts and the flowers
    Bracts in plantain (Plantago)
    NA
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bracts
    the flower has one or more bracts associated with it
    Flower bract length
    3 mm
    Hairs on underside of leaf
    the underside of the leaf is fuzzy or hairy
    Hairs on upper side of leaf
    the upper side of the leaf is not hairy, or it has very few hairs
    Hooked hairs on underside of leaf
    no
    Inflated hairs on leaf
    the leaf blade does not have inflated hairs on it
    Leaf arrangement
    opposite: there are two leaves per node along the stem
    Leaf blade base
    the leaf has a distinct leaf stalk (petiole)
    Leaf blade base symmetry
    the leaf blade base is symmetrical
    Leaf blade bloom
    the underside of the leaf has no noticeable bloom
    Leaf blade edges
    • the edge of the leaf blade has lobes, or it has both teeth and lobes
    • the edge of the leaf blade has teeth
    Leaf blade flatness
    the leaf is flat (planar) at the edges
    Leaf blade length
    30–80 mm
    Leaf blade primary vein pattern
    the secondary veins branch off at intervals from the primary vein
    Leaf blade shape
    • the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
    • the leaf blade is linear (very narrow with more or less parallel sides)
    • the leaf blade is oblong (rectangular but with rounded ends)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade texture
    the leaf blade is herbaceous (has a leafy texture)
    Leaf blade vein pattern
    the major veins of the leaf blade branch, but do not rejoin
    Leaf blade veins
    the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)
    Leaf blade width
    15–40 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf shiny
    the upper side of the leaf is dull or slightly shiny
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    the leaves have leaf stalks
    Leaf stalk attachment to leaf
    the petiole attaches at the basal margin of the leaf blade
    Leaf teeth and lobes
    • the leaf blade margin has forward-pointing teeth
    • the leaf has a row of two or more lobes on each side of the central axis
    Leaf tip
    • the tip of the leaf blade is acuminate (tapers to a long, thin point)
    • the tip of the leaf blade is acute (sharply pointed)
    Leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Leaf types
    NA
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaflet petiolules
    NA
    Leaves per node
    there are two leaves per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule edges
    NA
    Stipule features
    NA
    Stipule fused to leaf stalk
    NA
    Stipule length
    0 mm
    Stipule shape
    NA
    Stipules
    there are no stipules on the plant

  • Place

    Habitat
    • terrestrial
    • wetlands
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • edges of wetlands
    • shores of rivers or lakes
    • swamps

  • Scent

    Plant odor
    the plant does not have much of a smell

  • Stem, shoot, branch

    Branched tendrils
    NA
    Direction of stem hairs
    the hairs are pressed flat against the plant, pointing towards the plant's tip
    Flowering stem cross-section
    the flowering stem is roughly square
    Hair between stem nodes
    • the stem has hairs between the nodes
    • the stem has no hairs between the nodes
    Hairs between stem nodes
    the hairs on the stem are plain, without glands or branches, and not tangled
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    30–100 cm
    Stem bloom
    there is no powdery or waxy film on the stem
    Stem hair distribution
    the hairs on the stem are mainly found at the nodes
    Stem nodes swollen
    the stem is not swollen at the nodes
    Stem orientation
    the stems are upright or angled outwards
    Stem roughness between nodes
    the stem does not feel rough
    Stem succulence
    the stems are not succulent
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils
    Wings on stem
    the stem does not have wings on it

Wetland status

Occurs only in wetlands. (Wetland indicator code: OBL)

In New England

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
present
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

Massachusetts
widespread (S-rank: S5)

From Flora Novae Angliae dichotomous key

1.  Lycopus americanus Muhl. ex W. Bart. N

American water-horehound. Lycopus americanus Muhl. ex W. Bart. var. longii Benner; 
 L. americanus Muhl. ex W. Bart. var. scabrifolius Fern.; L. europaeus L. var. sinuatus (Ell.) Gray; L. pennsylvanicus Muhl. ex Raf.; L. pinnatifidus Raf.; L. sinuatus Ell.; Phytosalpinx americanus (Muhl.) Lunell • CT, MA, ME, NH, RI, VT; nearly throughout. Swamps, shorelines, wetland margins.

Native to North America?

Yes

Sometimes confused with

Lycopus uniflorus:
sepals triangular, shorter than 1 mm, as long as or shorter than the mature fruits, and leaf blades toothed (vs. L. americanus, with sepals narrow-triangular, longer than 1 mm, longer than the mature fruits, and leaf blades prominently toothed to lobed).
Lycopus virginicus:
sepals triangular, shorter than 1 mm, as long as or shorter than the mature fruits, and leaf blades toothed (vs. L. americanus, with sepals narrow-triangular, longer than 1 mm, longer than the mature fruits, and leaf blades prominently toothed to lobed).

Synonyms

  • Lycopus americanus var. longii Benner
  • Lycopus americanus var. scabrifolius Fern.
  • Lycopus europaeus var. sinuatus (Ell.) Gray
  • Lycopus pennsylvanicus Muhl. ex Raf.
  • Lycopus pinnatifidus Raf.
  • Lycopus sinuatus Ell.
  • Phytosalpinx americanus (Muhl.) Lunell

Family

Lamiaceae

Genus

Lycopus