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• All other flowering non-woody plants
• All other herbaceous, flowering dicots
• Lysimachia arvensis

Lysimachia arvensis — scarlet pimpernel

Facts

Scarlet pimpernel is an invasive species. The flowers close at the end of the first day of flowering, triggering self-fertilization in any flowers not already fertilized.

Habitat

Anthropogenic (man-made or disturbed habitats), brackish or salt marshes and flats, cliffs, balds, or ledges, coastal beaches (sea beaches), marshes, meadows and fields, ridges or ledges, wetland margins (edges of wetlands)

Characteristics

• Clonal plantlets

  Bulbils

  the plant does not appear to have bulbils

  Bulblets replace flowers

  there are no bulblets where the flowers are located

• Flowers

  Anther spurs

  the anthers do not have spurs on them

  Calyx growth after flowering

  the calyx does not grow to cover or partially cover the fruit

  Calyx symmetry

  there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)

  Carpels fused

  the carpels are fused to one another

  Cilia on petals

  the petal margins have cilia

  Cleistogamous flowers

  there are no cleistogamous flowers on the plan

  Corolla palate

  no

  Corona lobe length

  0 mm

  Epicalyx

  the flower does not have an epicalyx

  Epicalyx number of parts

  0

  Filament surface

  the filament has rough hairs or scales on it

  Flower appearance

  the flowers appear after the leaves have appeared

  Flower description

  the flower has a superior ovary, and lacks a hypanthium

  Flower diameter

  5–15 mm

  Flower orientation

  the flower points upwards or is angled outwards

  Flower petal color

  • blue to purple
  • orange
  • pink to red
  • white

  Flower reproductive parts

  the flower has both pollen- and seed-producing parts

  Flower symmetry

  there are two or more ways to evenly divide the flower (the flower is radially symmetrical)

  Flowers sunken into stem

  no

  Form of style

  • the style is knob-like at the tip, and unbranched
  • the style is narrow at the tip and unbranched

  Fringed petal edges

  • the petals are fringed
  • the petals are not fringed

  Fused stamen clusters

  there is one cluster of fused stamens

  Fusion of sepals and petals

  the petals or the sepals are fused into a cup or tube

  Horns in hoods (Asclepias)

  NA

  Hypanthium

  the flower does not have a hypanthium

  Hypanthium length

  0 mm

  Inflorescence one-sided

  the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks

  Inner tepals (Rumex)

  NA

  Interior flower disk

  the flower does not have an interior disc

  Length of flower stalk

  3–35 mm

  Marks on petals

  there are no noticeable marks on the petals

  Nectar spur

  the flower has no nectar spurs

  Number of branches in umbel

  0

  Number of carpels

  3–5

  Number of pistils

  1

  Number of sepals, petals or tepals

  there are five petals, sepals, or tepals in the flower

  Number of styles

  1

  Ovary position

  the ovary is above the point of petal and/or sepal attachment

  Perianth shape

  • the perianth is rotate (platter-shaped, the corolla flattened, circular, with nearly horizontally spreading lobes)
  • the perianth is salverform (with a slender basal tube and a more-or-less horizontally spreading mouth)

  Petal and sepal arrangement

  the flower includes two cycles of petal- or sepal-like structures

  Petal and sepal colors

  • blue to purple
  • orange
  • pink to red
  • white

  Petal appearance

  the petals are thin and delicate, and pigmented (colored other than green or brown)

  Petal folds or pleats

  the petals of the flower do not have folds or plaits

  Petal hairs (Viola)

  NA

  Petal length

  2–10 mm

  Petal length relative to sepals

  • the petals are about equal in length to the sepals
  • the petals are longer than the sepals

  Petal number

  5

  Petal shape

  the petal outline is obovate (roughly egg-shaped, but with the widest point above the middle of the leaf blade)

  Petal tip shape

  • the petal tip is lobed or fringed
  • the petal tip is obtuse (bluntly pointed)
  • the petal tip is rounded
  • the petal tip is truncate (ends abruptly in a more or less straight line as though cut off)

  Petal tips (Cuscuta)

  NA

  Raceme attachment (Veronica)

  NA

  Reproductive system

  all the flowers have both carpels and stamens (synoecious)

  Scales inside corolla

  no

  Sepal and petal color

  the sepals are different from the petals

  Sepal appearance

  the sepals are green or brown, and leaf-like in texture

  Sepal appendages

  the sepals do not have appendages on them

  Sepal appendages (Oenothera)

  NA

  Sepal auricles

  the sepals have no auricles

  Sepal cilia

  the sepals do not have cilia

  Sepal color

  green to brown

  Sepal length

  3.5–5 mm

  Sepal number

  5

  Sepal orientation

  the sepals are slightly curved outwards from the corolla

  Sepal relative length

  the sepal lobes are longer than the fused portion

  Sepal texture

  • the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
  • the sepals are thin and dry, paperlike

  Sepal tip shape

  • the sepal tip is acuminate (tapers to a very narrow point)
  • the sepal tip is acute (is sharply pointed)

  Sepal uniformity

  all the sepals are about the same size

  Sepals fused only to sepals

  the sepals are fused to each other (not other flower parts), at least near their bases

  Spur length

  0 mm

  Spur number

  NA

  Stamen attachment

  the stamens are attached at or near the bases of the petals or tepals

  Stamen lengths differ

  the stamens are all approximately the same length

  Stamen morphology

  the stamens within each cycle are the same

  Stamen number

  • 4
  • 5

  Stamen position relative to petals

  the stamens are lined up with the petals

  Stamens fused

  the stamens are attached to one another at or near their bases

  Staminodes

  there are no staminodes on the flower

  Stigma position

  the stigmas are positioned at the tip of the style

  Style petal-like

  the styles are not petal-like

  Surface of ovary

  the ovary surface has no points, bumps or wrinkles

  Umbel flower reproductive parts

  NA

  Upper lip of bilabiate corolla

  NA

• Fruits or seeds

  Achene relative orientation

  NA

  Achene shape

  NA

  Achene surface (Polygonum)

  NA

  Achene type

  NA

  Berry color

  NA

  Capsule color (Viola)

  NA

  Capsule ribs

  the capsule has no prominent ribs or wings

  Capsule splitting

  the capsule splits by two main valves, teeth or pores

  Carpel beak length

  0 mm

  Fruit (pyxis) dehiscence

  the pyxis splits on or near the middle

  Fruit cross-section

  the fruit is round in cross-section

  Fruit features (Brassicaceae)

  NA

  Fruit length

  3–6 mm

  Fruit length relative to sepals

  • the fruit is about the same length as its associated sepals
  • the fruit is longer than its associated sepals
  • the fruit is shorter than its associated sepals

  Fruit locules

  one

  Fruit shape

  the fruit is spherical

  Fruit stalk orientation

  the fruits curve or droop downwards

  Fruit type (general)

  the fruit is dry and splits open when ripe

  Fruit type (specific)

  the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)

  Hairs on fruit

  the fruits are not hairy

  Legumes (Fabaceae)

  NA

  Mericarp length

  0 mm

  Mericarp segment shape (Desmodium)

  NA

  Other markings on berry

  NA

  Ovary stipe

  the ovary or fruit does not have a stipe

  Ovary stipe length

  0 mm

  Placenta arrangement

  the plant has free-central placentation, where the ovules develop on a central column in a compound ovary lacking septa or with septa at the base only

  Rows of seeds in fruit (Brassicaceae)

  NA

  Schizocarpic fruit compression

  NA

  Schizocarpic fruit segments

  0

  Seed length

  1–1.5 mm

  Seed number

  12–45

  Seeds comose

  no hairs

  Septum in fruit (Brassicaceae)

  NA

  Wings on fruit

  the fruit does not have wings on it

  prickles on fruits

  the fruits do not have thorn-like defensive structures

• Glands or sap

  Glands on leaf blade

  the leaf blades have glandular dots or scales

  Sap

  the sap is clear and watery

  Sap color

  the sap is clear

• Growth form

  Growth form

  the plant is an herb (it has self-supporting stems)

  Horizontal rooting stem

  the plant does not have stolons

  Lifespan

  • the plant lives more than two years
  • the plant lives only a single year or less

  Parasitism

  the plant is not parasitic

  Plant color

  the leaves or young stems of the plant are green

  Plants darken when dry

  no

  Spines on plant

  the plant has no spines

• Leaves

  Bracteole edges

  NA

  Bracteole length

  0 mm

  Bracteole number (Apiaceae)

  0

  Bracteole shape

  NA

  Bracteoles

  there are no bracteoles on the plant

  Bracts in plantain (Plantago)

  NA

  Final leaf segment length (compound lvs only)

  0 mm

  Final leaf segment length to width ratio (compound lvs only)

  0

  Final leaf segment width (compound lvs only)

  0 mm

  Floral bracts

  the flower has one or more bracts associated with it

  Flower bract length

  Up to 30 mm

  Hairs on leaf stalk

  NA

  Hairs on underside of leaf

  the underside of the leaf is not hairy, or it has very few hairs

  Hooked hairs on underside of leaf

  no

  Inflated hairs on leaf

  the leaf blade does not have inflated hairs on it

  Leaf arrangement

  • opposite: there are two leaves per node along the stem
  • whorled: there are three or more leaves per node along the stem

  Leaf blade base

  • the leaf has no stalk
  • the leaf has no stalk and at the base it clasps the stem

  Leaf blade base symmetry

  the leaf blade base is symmetrical

  Leaf blade bloom

  the underside of the leaf has no noticeable bloom

  Leaf blade edges

  the edge of the leaf blade is entire (has no teeth or lobes)

  Leaf blade flatness

  • the edges of the leaf are curled under
  • the leaf is flat (planar) at the edges

  Leaf blade hairs

  NA

  Leaf blade length

  5–30 mm

  Leaf blade primary vein pattern

  the secondary veins branch off at intervals from the primary vein

  Leaf blade shape

  • the leaf blade is elliptic (widest near the middle and tapering at both ends)
  • the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
  • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)

  Leaf blade surface colors

  the upper side of the leaf blade is relatively uniform in color

  Leaf blade vein pattern

  the major veins of the leaf blade branch, but do not rejoin

  Leaf blade veins

  • the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)
  • the leaf blade has three or more main veins that start at or near the leaf blade base and run from the base towards the tip (it may or may not have secondary veins)

  Leaf blade width

  4–10 mm

  Leaf duration

  the leaves drop off in winter (or they whither but persist on the plant)

  Leaf form

  the leaves are green, with an expanded blade and a leaf-like texture

  Leaf hair orientation

  NA

  Leaf sheath length

  0 mm

  Leaf shiny

  the upper side of the leaf is dull or slightly shiny

  Leaf spines

  there are no spines on the leaf edges

  Leaf stalk

  the leaves have no leaf stalks, but attach directly to the stem

  Leaf stalk attachment to leaf

  NA

  Leaf stalk base

  NA

  Leaf stalk length

  0 mm

  Leaf teeth and lobes

  the edge of the leaf blade is entire (has no teeth or lobes)

  Leaf tip

  • the tip of the leaf blade is acute (sharply pointed)
  • the tip of the leaf blade is obtuse (bluntly pointed)

  Leaf tufts in axils

  there are no clusters of smaller leaves growing out of axils

  Leaf type

  the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)

  Leaf types

  NA

  Leaf variation

  the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem

  Leaflet number

  0

  Leaflet petiolules

  NA

  Leaves per node

  • there are five leaves per node along the stem
  • there are four leaves per node along the stem
  • there are three leaves per node along the stem
  • there are two leaves per node along the stem

  Pinnately compound leaf type

  NA

  Specific leaf type

  the leaves are simple (i.e., lobed or unlobed but not separated into leaflets)

  Stipels

  NA

  Stipule edges

  NA

  Stipule features

  NA

  Stipule fused to leaf stalk

  NA

  Stipule length

  0 mm

  Stipule shape

  NA

  Stipules

  there are no stipules on the plant

  Teeth per side of leaf blade

  0

• Place

  Habitat

  • terrestrial
  • wetlands

  New England state

  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont

  Specific habitat

  • brackish or salt marshes and flats
  • cliffs, balds, or ledges
  • edges of wetlands
  • man-made or disturbed habitats
  • marshes
  • meadows or fields
  • ridges or ledges
  • sea beaches

• Scent

  Plant odor

  the plant does not have much of a smell

• Stem, shoot, branch

  Branched tendrils

  NA

  Flowering stem cross-section

  the flowering stem is roughly square

  Hooked hairs on stem between nodes

  no

  Leaves on stem

  there is at least one full leaf above the base of the flowering stem

  Plant height

  6–50 cm

  Stem nodes swollen

  the stem is not swollen at the nodes

  Stem orientation

  the stems are upright or angled outwards

  Stem spacing

  the plant is solitary, or a few plants are growing together

  Stem succulence

  the stems are not succulent

  Tendril origin

  NA

  Tendrils

  the plant does not have tendrils

  Wings on stem

  the stem does not have wings on it

Wetland status

Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)

New England distribution and conservation status

From the dichotomous key of Flora Novae Angliae